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This subspecies is accepted, and its native range is Tropical Africa to KwaZulu-Natal, Madagascar, Arabian Peninsula.
Cyperus alternifolius subsp. flabelliformis

[FSOM]

M. Thulin et al. Flora of Somalia, Vol. 1-4 [updated 2008] https://plants.jstor.org/collection/FLOS

Morphology General Habit
Robust perennial with a 2–10 mm thick creeping woody rhizome and several stems usually placed in a straight row
Morphology Stem
Stems 25–150 cm long, 1.5–7 mm thick (but to 12 mm across the leaf-sheaths), rounded with longitudinal ridges, minutely scabrid; the basal part covered with leaf-sheaths and shorter black scales
Morphology Leaves
Leaf-blades absent or the uppermost sheath with a 1–8 cm long somewhat leafy limb (the function of the leaves has been taken over by the leafy involucral bracts)
Morphology Reproductive morphology Inflorescences
Inflorescence a compound anthela 5–25 cm in diam., subtended by 15–25 leafy involucral bracts; bracts 10–35 cm long and 3–20 mm wide, scabrid on margin and major ribs, spirally arranged along a 1–5 cm long axis; largest peduncle 3–13 cm long; each spikelet-cluster digitate consisting of 3–20 spikelets, if more than 10 then the cluster is almost capitate
Morphology Reproductive morphology Inflorescences Spikelets
Spikelets 2–10 x 1–2.5 mm, ligth to medium brown, lanceolate, much compressed, 15–30-flowered
Morphology Reproductive morphology Inflorescences Bracts Glume
Glumes 1–2 mm long, straw-coloured or golden to reddish brown with 3-keeled midrib excurrent in a short mucro or acute tip
Morphology Reproductive morphology Flowers Gynoecium Style
Style 3-branched
Morphology Reproductive morphology Fruits
Nutlet 0.8–0.9 x 0.4–0.5 mm, triangular, lanceolate to oblong, yellow as young turning brownish when mature, minutely papillose.
Distribution
N1 widespread in tropical Africa.
Ecology
Altitude range 1000–1300 m.

[FTEA]

Cyperaceae, K Hoenselaar, B. Verdcourt & H. Beentje. Hypolytrum, D Simpson. Fuirena, M Muasya. Flora of Tropical East Africa. 2010

Type
Type: Ethiopia, Adua, Schimper 55 (P, holo.; HAL, K!, iso.)
Morphology General Habit
Perennial, robust, up to 2 m tall, with a creeping rhizome, 2–10 mm in diameter and several culms usually placed in a row.
Morphology Culms
Culms 66–160 cm long, 3–8 mm wide, rounded to trigonous, with longitudinal ridges, glabrous to sometimes minutely scabrid, the base covered with black fibrous remains from old leaf sheaths.
Morphology Leaves Leaf sheaths
Sheath almost black, pale brown to green, 1–36 cm long; blade absent
Morphology Reproductive morphology Inflorescences Bracts
Involucral bracts many. Leaf-like, spreading, spirally arranged along a 1–5 cm long axis, 18–37 cm long, 0.8–1.5 cm wide, linear, flat, scabrid, apex acute
Morphology Reproductive morphology Inflorescences
Inflorescence compound, primary branches many, 3.5–10 cm long.
Morphology Reproductive morphology Inflorescences Spikelets
Spikelets in digitate clusters at the end of secondary and tertiary branches, 5–20 per cluster, lanceolate to elliptic-ovoid, much compressed, 3.5–11 mm long, 1.4–3 mm wide.
Morphology Reproductive morphology Inflorescences Bracts Glume
Glumes very pale brown, golden brown to reddish brown, elliptic-ovate, 1.4–2.2 mm long, 0.9–1.6 mm wide, 3-keeled, green, apex acute, sometimes slightly mucronate, glabrous
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 3: filaments 1.6–2.2 mm long.
Morphology Reproductive morphology Flowers Androecium Stamens Anthers
Anthers 0.8–1.4 mm long; the connective protruding into a needlelike apex
Morphology Reproductive morphology Fruits
Nutlet yellow to brown, narrowly ovoid to oblong, trigonous, 0.7–1.1 mm long, 0.5–0.6 mm wide, minutely papillose
Figures
Fig. 28, p 188
Ecology
Swamps, grasslands, lake shores, streambanks; 30–2150 m
Conservation
Least Concern (LC) due to its wide distribution and common habitat.
Note
Widely distributed as an ornamental.
Distribution
Range: Senegal, Guinea, Sierra Leone, Liberia, Ghana, Nigeria, Cameroon, Congo Brazaville, Congo-Kinshasha, Rwanda, Burundi, Sudan, Ethiopia, Somalia, Angola, Zambia, Malawi, Zimbabwe, South Africa Flora districts: U1 U2 U3 U4 K1 K2 K4 K6 K7 T1 T2 T3 T4 T5 T6 T7 T8 Z

[CPLC]

Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

Distribution
Naturalizada en Colombia; Alt. 900 - 2900 m.; Andes, Valle del Cauca.
Morphology General Habit
Hierba, acuática
Conservation
No Evaluada

Native to:

Angola, Benin, Botswana, Burundi, Cameroon, Chad, Congo, Djibouti, Eritrea, Ethiopia, Ghana, Guinea, Kenya, KwaZulu-Natal, Liberia, Madagascar, Malawi, Mauritania, Mozambique, Nigeria, Rwanda, Saudi Arabia, Senegal, Sierra Leone, Somalia, Sudan, Tanzania, Uganda, Yemen, Zambia, Zaïre, Zimbabwe

Introduced into:

Algeria, Azores, Bahamas, Baleares, Bangladesh, Bermuda, California, Canary Is., Cape Verde, Caroline Is., Chile North, China Southeast, Colombia, Comoros, Cook Is., Corse, Costa Rica, Cuba, Dominican Republic, East Aegean Is., East Himalaya, Egypt, Fiji, Florida, France, Galápagos, Haiti, Hawaii, Honduras, Italy, Jamaica, Jawa, Leeward Is., Louisiana, Madeira, Marquesas, Mauritius, Mexico Central, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Myanmar, Nansei-shoto, New Caledonia, New York, New Zealand North, Norfolk Is., Ogasawara-shoto, Pakistan, Palestine, Philippines, Portugal, Puerto Rico, Réunion, Sardegna, Society Is., Spain, Sri Lanka, St.Helena, Sumatera, Taiwan, Texas, Thailand, Trinidad-Tobago, Tunisia, Tuvalu, Venezuela, Venezuelan Antilles, Windward Is.

Cyperus alternifolius subsp. flabelliformis Kük. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Schimper [55], Ethiopia K000321368 isotype

First published in H.G.A.Engler (ed.), Pflanzenr., IV, 20(101): 193 (1936)

Accepted by

  • Boulos, L. (2005). Flora of Egypt 4: 1-617. Al Hadara Publishing, Cairo.
  • Govaerts, R. & Simpson, D.A. (2007). World Checklist of Cyperaceae. Sedges: 1-765. The Board of Trustees of the Royal Botanic Gardens, Kew.
  • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
  • Smith, A.C. (1979). Flora Vitiensis Nova. A new flora for Fiji (Spermatophytes only) 1: 1-495. Pacific Tropical Botanical Garden, Lawai.
  • Tanaka, N., Koyama, T. & Murata, J. (2005). The flowering plants of Mt. Popa, central Myanmar - Results of Myanmar-Japanese joint expeditions, 2000-2004 Makinoa 5: 1-102.
  • Verloove, F. (2014). A conspectus of Cyperus s.l. (Cyperaceae) in Europe (incl. Azores, Madeira and Canary islands), with emphasis on non-native naturalised species Webbia; Raccolta de Scritti Botanici 69: 179-223.

Not accepted by

  • Hoenselaar, K., Verdcourt, B. & Beentje, H. (2010). Cyperaceae Flora of Tropical East Africa: 1-466. [Cited as Cyperus involucratus.]
  • Wu, Z. & Raven, P.H. (eds.) (2010). Flora of China 23: 1-515. Missouri Botanical Garden Press, St. Louis. [Cited as Cyperus involucratus.]

Literature

Kew Backbone Distributions

  • Acevedo-Rodríguez, P. & Strong, M.T. (2005). Monocotyledons and Gymnosperms of Puerto Rico and the Virgin Islands Contributions from the United States National Herbarium 52: 1-415.
  • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
  • Audru, J., Cesar, J. & Lebrun, J.-P. (1993). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 2(2): 433-968. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
  • Barthelat, F. (2019). La flore illustrée de Mayotte: 1-687. Biotope éditions.
  • Bernal, R., Gradstein, R.S. & Celis, M. (eds.) (2016). Catálogo de plantas y líquenes de Colombia 1-2: 1-3068. Libro impreso.
  • Browning, J. & al. (2020). Flora Zambesiaca 14: 1-455. Royal Botanic Gardens, Kew.
  • Brundu, G. & Camarda, I. (2013). The Flora of Chad: a checklist and brief analysis PhytoKeys 23: 1-18.
  • Castroviejo, S. & al. (eds.) (2008). Flora Iberica 18: 1-420. Real Jardín Botánico, CSIC, Madrid.
  • Cossu, T.A, Camarda, I. & Brundu, G. (2014). A catalogue of non-native weeds in irrigated crops in Sardinia (Italy) Webbia; Raccolta de Scritti Botanici 69: 145-156.
  • Dobignard, D. & Chatelain, C. (2010). Index synonymique de la flore d'Afrique du nord 1: 1-455. Éditions des conservatoire et jardin botaniques, Genève.
  • Dufour-Dror, J.M. & Fragman-Sapir, O. in Dufour-Dror, J.M. (2019). Alien Plant Species in Natural & Disturbed Areas in Israel Alien Invasive Plants in Israel, ed. 2: 1-246. Dan Perry Pub., Nature & Parks Authority, Israel Ministry of Environmental Protection.
  • Hammel, B.E. & al. (2003). Manual de Plantas de Costa Rica 2: 1-694. Missouri Botanical Garden Press, St. Louis.
  • Hassler, M. (2012). Flora of Rhodes. Systematic list of flora of Rhodes http://www.rz.uni-karlsruhe.de/~db111/flora/rhodos/list.php.
  • Hoenselaar, K., Verdcourt, B. & Beentje, H. (2010). Cyperaceae Flora of Tropical East Africa: 1-466.
  • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
  • Jaramillo Díaz, P. & Guézou, A. (2017). CDF Checklist of Galapagos Vascular Plants - FCD Lista de especies de Plantas Vasculares de Galápagos http://www.darwinfoundation.org/datazone/checklists/vascular-plants/.
  • Kitalong, A.H., DeMeo, R.A. & Holm, T. (2008). Native Trees Palau. A field guide: 1-236.
  • Lambdon, P. (2012). Flowering plants & ferns of St Helena: 1-624. Pisces publications for St Helena nature conservation group.
  • Lorence, D.H. & Wagner, W.L. (2019). Flora of the Marquesas Islands 1: 1-411. National Tropical Botanic Garden, Smithsonian, DRPF.
  • MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
  • Marais, W. & Araujo, A.C. (2018). Flore des Mascareignes 202: 1-99. IRD Éditions, MSIRI, RBG-Kew, Paris.
  • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
  • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
  • Tanaka, N., Koyama, T. & Murata, J. (2005). The flowering plants of Mt. Popa, central Myanmar - Results of Myanmar-Japanese joint expeditions, 2000-2004 Makinoa 5: 1-102.
  • Verloove, F. (2014). A conspectus of Cyperus s.l. (Cyperaceae) in Europe (incl. Azores, Madeira and Canary islands), with emphasis on non-native naturalised species Webbia; Raccolta de Scritti Botanici 69: 179-223.
  • Werier, D. (2017). Catalogue of the Vascular plants of New York state Memoirs of the Torrey Botanical Club 27: 1-542. New York Botanical Garden.
  • Woodroffe, C.D. (1985). Vegetation and flora of Nui Atoll, Tuvalu Atoll Research Bulletin 283: 1-18.

Flora of Somalia

  • Cyp. E. Afr.: 4 (1936).
  • Flora Somalia, Vol 4, (1995) Author: by K. A. Lye [updated by M. Thulin 2008]

Catálogo de Plantas y Líquenes de Colombia
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Somalia
Flora of Somalia
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Tropical East Africa
Flora of Tropical East Africa
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Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Science Photographs
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/