Skip to main content
This species is accepted, and its native range is Bolivia.
A specimen from Kew's Herbarium


Wood, J.R.I. Kew Bull (2011) 66: 199.

A Bolivian endemic only known from the western side of the Cuesta de Sama near Tarija where it grows on open grassy slopes on stony soil between 2800 – 3200 m.
Morphology General Habit
Low, untidy, much-branched aromatic undershrub to 30 cm in height; stems quadrangular, pilose, the leaf nodes scurfy-pilose
Morphology Leaves
Leaves petiolate, very crowded; petioles 1 – 1.5 mm long, pilose; lamina 7 – 12 × 3 – 6 mm, ovate, obtuse, cuneate at the base, margin obscurely 2 – 3-toothed on each side, both surfaces pilose with large-celled hairs, the lower surface paler and very densely dotted with pale glands
Morphology Reproductive morphology Fruits
Nutlets 1 – 1.5 × 1 mm, obscurely angled, obtuse, pubescent with stiffish hairs.
Morphology Reproductive morphology Inflorescences
Inflorescence of 1 – 3-flowered axillary cymes; peduncle c. 1 mm long, pilose; bracts at branching point, c. 2 × 1 mm, shorter than the pedicels, oblong to narrowly oblong-elliptic, pilose; pedicels 2 – 3 mm long, pilose, not conspicuously jointed above the base; calyx 2 – 2.5 mm long, narrowly campanulate, ribbed, the exterior pilose, gland-dotted, the interior glabrous except for a ring of hairs around the base of the teeth, teeth 1 – 1.5 × 1 mm, longer than the tube, lanceolate, acuminate, the tips recurved; corolla white, subequally 5-lobed, the tube c. 1 mm long, ± equalling the calyx tube; filaments very short, subequal, anthers included; style c. 1 mm long, glabrous, shortly exserted
Obviously related to both Clinopodium cochabambense and C. odorum but distinguished by the obtuse nutlets and the pilose indumentum of large-celled hairs, which cover all vegetative parts. Although these two characters both suggest some affinity with C. gilliesii which grows within a few kilometres of the type locality on the western side of the Cuesta de Sama, this species does not appear to be of hybrid origin. The populations are not mixed, C. pilosum is perfectly fertile and the hairy nutlets, ovate leaves and the shape of the calyx teeth all place it close to C. odorum rather than C. gilliesii.

Native to:


Clinopodium pilosum J.R.I.Wood appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Wood, J.R.I. [21385], Bolivia K000975976 holotype
Wood, J.R.I. [21385], Bolivia K000975975 holotype

First published in Kew Bull. 66: 213 (2011)

Accepted by

  • Govaerts, R., Nic Lughadha, E., Black, N., Turner, R. & Paton, A. (2021). The World Checklist of Vascular Plants, a continuously updated resource for exploring global plant diversity. Scientific Data 8: 215.


Kew Bulletin

  • Bell, C. D. & Donoghue, M. J. (2005). Phylogeny and biogeography of Valerianaceae (Dipsacales) with special reference to the South American valerians. Organ. Divers. Evol. 5: 147 – 159.CrossRefGoogle Scholar
  • Bentham, G. (1832 – 5). Labiatarum Genera et Species. London, 1 – 60 (1832), 61 – 322 (1833), 323 – 645 (1834), 646 – 783 (1835).
  • Briquet, J. (1896). Fragmenta Monographiae Labiatarum. Bull. Herb. Boiss. 4: 762 – 878.
  • Briquet, J. (1897). Labiatae. In: A. Engler & K. Prantl, Die Natürlichen Pflanzenfamilien IV(3): 183 – 373. Wilhelm Englemann, Leipzig.
  • Cantino, P. D. & Wagstaff, S. J. (1998). A Reexamination of North American Satureja s.l. (Lamiaceae) in light of molecular evidence. Brittonia 50: 63 – 70.CrossRefGoogle Scholar
  • Epling, C. & Játiva, C. (1964). Revisión del genero Satureja en América del Sur. Brittonia 16: 393 – 416.CrossRefGoogle Scholar
  • Epling, C. (1927). Studies in South American Labiatae III. Ann. Missori Bot. Gard. 14: 47 – 86.CrossRefGoogle Scholar
  • Epling, C. (1937). Synopsis of South American Labiatae. Repert. Spec. Nov. Regni Veg. 85: 97 – 192.
  • Fries, R. E. (1905). Zur Kenntnis der alpinen Flora in nördlichen Argentinien. Nov. Acta Regiae Soc. Sci Upsal. Ser. 4: 1 – 205.
  • Govaerts, R. (1999). World Checklist of Seed Plants 3(1). Continental Publishing, Deurne.
  • Grisebach, A. H. R. (1874). Plantae Lorentzianae. Abh. Königl. Ges. Wiss. Göttingen 19: 49 – 279.
  • Grisebach, A. H. R. (1879). Symbolae ad Floram Argentiniam. Abh. Königl. Ges. Wiss. Göttingen 24 (1): 1 – 345.
  • Harley, R. M. & Granda Paucar, A. (2000). List of species of tropical American Clinopodium (Labiatae) with new combinations. Kew Bull. 55: 917 – 927.CrossRefGoogle Scholar
  • Hieronymus, G. H. E. W. (1882). Plantae diaphoricae florae argentiniae. Bol. Acad. Nac. Ci. Cordova 4: 182 – 598.
  • Hughes, C. E. & Eastwood, R. J. (2006). Island Radiation on a continental scale: exceptional rates of plant diversification after uplift of the Andes. Proc. Nat. Acad. Sci. 103: 10334 – 10339.PubMedCrossRefGoogle Scholar
  • IUCN (2001). IUCN Red List Categories and Criteria: Version 3.1. Prepared by the IUCN Species Survival Commission. IUCN, Gland, Switzerland & Cambridge, UK.
  • Kuntze, C. E. O. (1891 – 1898). Revisio Generum Plantarum 2: 515 (1891); 3: 259 (1898). A. Felix, Leipzig.
  • Leybold, F. (1873). Excursión a las Pampas Argentinas. Imprenta Nacional, Santiago.
  • Pontiroli, A. (1993). Labiatae. In: A. L. Cabrera, Flora de la Provincia de Jujuy IX, Verbénaceas a Caliceraceas. INTA, Buenos Aires.
  • Rusby, H. H. (1896). An enumeration of the plants collected in Bolivia by Miguel Bang [3]. Mem. Torrey. Bot. Club 6: 1 – 130.
  • Schmidt-Lebuhn, A. N. (2008). A Revision of the Genus Minthostachys (Labiatae). Mem. New York Bot. Gard. 98: 1 – 75.
  • Torrico, G., Peca, C., Beck, S. G. & García, E. (1994). Leñosas Utiles de Potosí. Potosí.
  • Wagstaff, S. J., Olmstead, R. G. & Cantino, P. D. (1995). Parsimony analysis of cpDNA restriction site variation in subfamily Nepetoideae (Labiatae). Amer. J. Bot. 82: 886 – 892.CrossRefGoogle Scholar
  • Weddell, A. A. (1857). Chloris Andina. Vol. 2. P. Bertrand, Paris.
  • Xifreda, C. C. (1999). Lamiaceae. In: O. Zuloaga & O. Morrone (eds), Catálogo de las Plantas Vasculares de la Republica Argentina II, Angiospermae (Dicotyledoneae). Monogr. Syst. Bot. Missouri Bot. Gard. 74: 769 – 781.

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 World Checklist of Selected Plant Families.

Kew Bulletin
Kew Bulletin

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.