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This species is accepted, and its native range is SW. Cameroon.
A specimen from Kew's Herbarium


Cheek, M. & Etuge, M. 2009. Allophylus conraui (Sapindaceae) reassessed and Allophylus ujori described from Cameroon Highlands of West Africa. Kew Bulletin 64: 495. DOI:

Conservation Status . Since ‘Kebo’, the original type locality of Allophylusconraui has not been refound, only four of the five localities for the species are known. Of the eight specimens collected, five derive from two localities in the Bakossi Mts. Three of these specimens were collected at about 1000 – 1100 m alt. on a path to the summit of Mt Kupe that begins between the villages of Nyasoso and Mbulle — this site has the prospect of being protected in the future since the Bakossi support proposed government protection of the forest on Mt Kupe as an Integrated Ecological Rseserve (Cheek et al. 2004). Two specimens are known from the lowlands of W BakossiMts between Banyemem and Ayon. This site is vulnerable to unregulated logging. A little to the N, in the BanyangMbo Reserve, at similar (c. 300 m) alt., a sixth specimen is known, in a relatively secure site. The fourth site, newly discovered in 2006, at Bechati in the Lebialem Highlands, is under threat of forest clearance for agriculture. An area of occupancy of 20 km² is estimated here for the species, based on a grid cell size of 4 km². Accordingly A. conraui is here assessed as Endangered EN B2a,b(iii) under IUCN (2001).
South West Province, Cameroon.
Lowland and submontane evergreen forest; 200 – 1100 m.
Disc lobes 4, asymmetrically inserted, rounded-hexagonal, c. 0.5 × 0.5 mm, outer surface with c. 6 hairs, inner surface with 10 – 15 reflexed hairs
Morphology General Habit
Shrub 0.8 – 1.5 m tall, unarmed, main stem dark brown, slightly and finely longitudinally grooved, thinly scattered with inconspicuous circular orange, raised lenticels of c. 0.5 mm diam., glabrous; leafy branches densely white or yellowish-white pilose, hairs simple, 3 mm long
Morphology Leaves
Leaves membranous, drying dark brown or green above, grey-green below, with three subequal, subsessile leaflets, lateral leaflets oblanceolate-elliptic, 9.4 – 13 × 3.5 – 5 cm, acumen c. 1 cm long, slender, acute, base acute, margin coarsely serrate, (4 –) 6 – 14 teeth per side, teeth 1.5 – 2 (– 3) mm deep, 3 – 5 mm wide, terminating the secondary nerves, upper and lower surface pilose, densely so along the midrib, hairs as the leafy branches, subepidermal layer of lower surface with cells frequently bearing a single isodiametric crystal of c. 0.01 mm diam. (visible with light microscope after clearing), domatia absent, petiolule absent; central leaflet slightly broader than the laterals, base cuneate, petiolule 0.15 – 0.5 cm, pilose; petiole terete, 3.5 – 5.5 (– 8) cm long, white pilose
Morphology Reproductive morphology Flowers
Female flowers as the male but disc lobes fleshier Male flower buds near anthesis to 1 × 1.3 mm, flowers at anthesis white
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 8, erect, filaments 0.6 – 1.15 mm long, tapering to the apex, sparsely long-crinkled hairy, hairs 0.1 – 0.15 mm long; anthers oblong, 0.4 – 0.5 mm long, apex slightly retuse, base deeply sagittate; pistillode 0.1 – 0.3 mm long, inconspicuous
Morphology Reproductive morphology Flowers Calyx
Inner sepals 2, tranversely elliptic, deeply concave, 1 × 1.2 mm, 1-nerved, with 1 or 2 hairs on the outer surface Outer sepals 2, obovate, strongly concave, c. 1 × 0.6 mm, outer surface with 6 – 10 hairs each 0.2 – 0.3 mm long
Morphology Reproductive morphology Flowers Corolla
Petals 4, obovate-obtriangular, c. 0.7 × 0.5 mm, apex rounded, base cuneate, abaxial surface glabrous apart from two tufts of hairs each 0.1 – 0.2 mm long, projecting from the sides at the middle, adaxial surface with a densely hairy ligule occupying the lower two-thirds, upper third glabrous
Morphology Reproductive morphology Flowers Gynoecium Carpels
Carpels separated by the style base, globose, 0.8 – 1 × 0.6 – 0.7 × c. 0.4 mm, densely white hairy, hairs to 0.5 mm long
Morphology Reproductive morphology Flowers Gynoecium Pistillodes
Pistillode c. 2 mm long, bicarpellate, apocarpous, style bifid
Morphology Reproductive morphology Flowers Gynoecium Style
Style gynobasic, free, united part 0.5 – 0.6 mm long; stylar arms separated by 90 – 180°, 0.8 – 1 mm long, inner surface shortly papillate-hairy, presumably stigmatic. Infructescence not accrescent, 1 – 4-fruited
Morphology Reproductive morphology Fruits
Fruit red, obovoid, 8 – 11 × 7 – 8 mm, appressed-puberulent, single-seeded
Morphology Reproductive morphology Inflorescences
Inflorescence axillary, single, 1 or 2 per leafy branch, unbranched, 2 – 4 cm long, sparingly white pilose, hairs up to 0.7 mm long; peduncle 0.3 – 0.4 (– 1) cm long; cymules 20 – 35; bracts basal, linear, c. 0.5 mm long; cymule-peduncles c. 1 mm long, rhachis 1 – 2 mm long, 1 – 3-flowered, bracteoles at the base of the pedicels, c. 0.3 mm long; pedicel c. 1.75 mm long, articulated midway along its length
Morphology Reproductive morphology Seeds
Seed ovoid, c. 7 × 5 mm
Notes . The exact location, ‘Kebo’, of the collection site of the original specimen, Conrau 219 is not clear. It has not been found in the list of Cameroonian botanical collection sites (Letouzey1968) nor in the US Board on Geographic Names (1962), nor in Google map. Consultation with our Cameroonian colleague Bate Oben, who works on georeferencing of Cameroonian specimens, has also not helped find the site. One suggestion is that Kebo is a Germanic version of the modern Batibo with the usual Bantu place name prefix Ba- omitted, but this idea needs more support. It is unlikely that ‘Kebo’ is a typographical error since it has been applied elsewhere. For example the location of Conrau 214, a specimen of Allophylusgrandifolius (Baker) Radlk., a lowland species, cited by Keay (1958: 714) is also given as Kebo. Conrau collected 276 numbers in then German Kamerun in 1898 and 1899 (Hepper & Neate1973). Other Conrau collection sites in Cameroon are confined to a fairly small area extending from the Bamiliké-BamboutosMts (e.g. Bangwa) of modern W Province, to Bali, further north, in the Bamenda Highlands of contemporary NW Province, so it is likely that Kebo falls within this area. Most of the specimens of the taxon we here elect to name A. conraui derive from the Bakossi area, to the S of Conrau’s collection sites. The exception is Tchiengue 2787 from Bechati, in the general vicinity of Conrau’sBangwa collection site, and for this reason this specimen has been chosen as neotype here. Etymology . Named for Gustav Conrau (? – 1899) who, apart from collecting the specimen on which Allophylusconraui was based, collected about 250 specimen numbers in western Cameroon when working with the Victoria Plantation Company (Hepper & Neate1973). Allophylus sp. 1, Cheek & Etuge (2004: 398, fig. 13D).
Probably western Cameroon, Kebo (not located exactly by us), fl., immature fruit, June 1899, (holotype B†); Lebaliem Highlands, Bechati, 320 m alt., fl. buds, fr. 25 Sept. 2006, Tchiengue 2787 (neotype K, selected here; isoneotypes WAG, YA).


Sapindaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:2. 1958

Morphology General Habit
A shrub
Morphology Reproductive morphology Flowers
White flowers
In forest.

Local Names & Uses .NyongNyong (Bakossi-Muetug) used to treat leprosy in combination with two other herbs fide EpieNgome — Gosline 137).

Native to:


Allophylus conraui Gilg ex Radlk. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Dec 1, 2007 Olorunfemi, J. [FHI 30611], Cameroon K000093181
Dec 1, 2007 Letouzey, R. [13754], Cameroon K000093178
Dec 1, 2007 Cable, S. [3327], Cameroon K000197912
Dec 1, 2007 Gosline, G. [137], Cameroon K000197911
Dec 1, 2007 Cheek, M. [10574], Cameroon K000510704
Jan 1, 2007 Tchiengue, B. [2787], Cameroon K000510701 Unknown type material
Jan 1, 2007 Tchiengue, B. [2787], Cameroon K000461994 Unknown type material
Nov 1, 2003 Cable, S. [3559], Cameroon K000197913

First published in Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 38: 221, 237 (1908 publ. 1909)

Accepted by

  • Govaerts, R. (1995). World Checklist of Seed Plants 1(1, 2): 1-483, 529. MIM, Deurne.


Flora of West Tropical Africa

  • Pflanzenr. 529.
  • in Sitzb. Math -Phys. Akad. Münch. 28: 221 (1908)

Flora of West Tropical Africa
Flora of West Tropical Africa

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Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
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