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This species is accepted, and its native range is Mexico.
Physalis ixocarpa

[FZ]

Flora Zambesiaca. Vol. 8, Part 4. Solanaceae. Gonçalves AE. 2005

Morphology General
Erect, occasionally spreading or trailing, weak to ± robust, ± dichotomously branched, annual herb, (7)20–65 cm high (elsewhere said to reach 1. 3 m), yellowish-green or green, sometimes tinged purple, ± sparsely clothed with simple, ± appressed to patent, eglandular, occasionally short and minute, glandular hairs, also relatively long glandular hairs, furnished with ± sessile glands too
Morphology Branches
Branches subterete or angular, striate, drying ± sulcate, ± scabrous-pubescent or subglabrous Branches subterete or angular, striate, drying ± sulcate, ± scabrous-pubescent or subglabrous.
Morphology Leaves
Leaves usually solitary; petiole (0.3)1–4(4.5) cm long, often slightly winged, rather sheathing at the base; lamina membranous or ± fleshy, (1)2–11 × (0.5)1–5.5 cm, ovate or ovate-lanceolate, occasionally ovate-deltate, rhombic, elliptic or oblong, base obtuse, cuneate or attenuate, and often unequal-sided or dimidiate, decurrent into the petiole, apex acute, occasionally sub-acute or ± acuminate, sinuate-dentate or dentate, the teeth unequal, rather obtuse or rarely sub-acute, the sinuses ± deep, obtuse, glabrous except where decurrent on the petiole, sometimes with a few, often short, appressed hairs on the nerves and near the margins, occasionally scattered elsewhere Leaves usually solitary; petiole (0.3)1–4(4.5) cm long, often slightly winged, rather sheathing at the base; lamina membranous or ± fleshy, (1)2–11 × (0.5)1–5.5 cm, ovate or ovate-lanceolate, occasionally ovate-deltate, rhombic, elliptic or oblong, base obtuse, cuneate or attenuate, and often unequal-sided or dimidiate, decurrent into the petiole, apex acute, occasionally sub-acute or ± acuminate, sinuate-dentate or dentate, the teeth unequal, rather obtuse or rarely sub-acute, the sinuses ± deep, obtuse, glabrous except where decurrent on the petiole, sometimes with a few, often short, appressed hairs on the nerves and near the margins, occasionally scattered elsewhere.
Morphology Reproductive morphology Flowers
Flowers solitary, axillary, erect to pendulous. Flowers solitary, axillary, erect to pendulous; pedicel greenish or violaceous, (3)4–6(7) mm long, ± hairy and glandular, in fruit elongating to 8(10) mm.
Morphology Reproductive morphology Flowers Pedicel
Pedicel greenish or violaceous, (3)4–6(7) mm long, ± hairy and glandular, in fruit elongating to 8(10) mm
Morphology Reproductive morphology Flowers Calyx
Calyx (3.5)4–5.5(6) mm long, 2.5–3.5 mm across at the base of the lobes, campanulate, sub-angled or ribbed, truncate or invaginated at the base, puberulous mostly along the nerves, usually furnished also with ± sessile glands, more abundant towards the base, on the inside glabrous except for the lobes with ± dense, minute indumentum especially towards the apex and near the margins; lobes unequal or subequal, 0.8–2 × 0.6–1. 8 mm, ± triangular, obtuse or ± acute, occasionally bluntly acuminate; in fruit green or yellowish-green, sometimes purple towards the base or also with purple venation, 18–32 × (15)17–25 mm, ovoid or subglobose, ± terete or rather ribbed, sub-acute or bluntly sub-acuminate, half to three-quarters filled by the fruit, ± glabrescent, the lobes 2.4–4 × 2–3.5 mm Calyx (3.5)4–5.5(6) mm long, 2.5–3.5 mm across at the base of the lobes, campanulate, sub-angled or ribbed, truncate or invaginated at the base, puberulous mostly along the nerves, usually furnished also with ± sessile glands, more abundant towards the base, on the inside glabrous except for the lobes with ± dense, minute indumentum especially towards the apex and near the margins; lobes unequal or subequal, 0.8–2 × 0.6–1.8 mm, ± triangular, obtuse or ± acute, occasionally bluntly acuminate; in fruit green or yellowish-green, sometimes purple towards the base or also with purple venation, 18–32 × (15)17–25 mm, ovoid or subglobose, ± terete or rather ribbed, sub-acute or bluntly sub-acuminate, half to three-quarters filled by the fruit, ± glabrescent, the lobes 2.4–4 × 2–3.5 mm.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens usually exserted, subequal; filaments violaceous, (1)1. 2–2.5 mm long, filiform, attached to the corolla tube near the base, sometimes furnished with a few hairs; anthers violaceous or bluish-violet, (1)1. 2–1. 7(2) mm long, ovate-oblong to elliptic in outline, straight or slightly bent after anthesis Stamens usually exserted, subequal; filaments violaceous, (1)1.2–2.5 mm long, filiform, attached to the corolla tube near the base, sometimes furnished with a few hairs; anthers violaceous or bluish-violet, (1)1.2–1.7(2) mm long, ovate-oblong to elliptic in outline, straight or slightly bent after anthesis.
Morphology Reproductive morphology Flowers Disc
Disk 0.1–0.2 mm high, fleshy, glabrous Disk 0.1–0.2 mm high, fleshy, glabrous.
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary 1. 2–2 × 0.8–1. 5 mm, ovoid or subglobose, glabrous.
Morphology Reproductive morphology Flowers Gynoecium Style
Style greenish-blue, (1)1. 5–2.5(3) mm long, filiform, straight or curved upwards
Morphology Reproductive morphology Fruits
Fruit green to yellowish, pale violet or violaceous, viscid, ± sessile on the invaginated base of the erect to pendulous calyx, 10–13 mm in diameter, subglobose Fruit green to yellowish, pale violet or violaceous, viscid, ± sessile on the invaginated base of the erect to pendulous calyx, 10–13 mm in diameter, subglobose.
Note
Chromosome number: 2n=24 Common name: “Wild Cape Gooseberry”.
Ecology
Cleared bushland, seasonally flooded grassland and cultivated ground, on roadsides and anthills, sometimes as a weed of cultivation; up to 1400 m.
Distribution
Botswana. Zimbabwe BOT SE, ZAM C, ZAM S, ZIM W, ZIM C, ZIM E, ZIM S Origin unknown, naturalized in Portugal and probably also in Holland and Australia. In Africa recorded also from Dem. Rep. Congo. Zambia
Morphology General Habit
Erect, occasionally spreading or trailing, weak to ± robust, ± dichotomously branched, annual herb, (7)20–65 cm high (elsewhere said to reach 1.3 m), yellowish-green or green, sometimes tinged purple, ± sparsely clothed with simple, ± appressed to patent, eglandular, occasionally short and minute, glandular hairs, also relatively long glandular hairs, furnished with ± sessile glands too.
Morphology Reproductive morphology Flowers Corolla
Corolla yellow or greenish-yellow, blotched with 5 dark brownish violet or purplish markings strongly contrasting with the surrounding limb, sometimes spreading downwards and somewhat along the nerves upwards, 4–7(8) mm long, rotate; tube subglabrous, furnished with few ± sessile glands, on the inside with felted indumentum near the insertion of the stamens and with a ring of hyaline hairs at the mouth; limb (5)6–9 mm across, shortly 5-lobed, erect to widely spreading, sometimes reflexed when fully expanded, on the outside with ± appressed, short hairs and also sessile glands often along the nerves, on the median region of each lobe on the part not folded in bud, glabrous inside except for the lobes with minute hairs towards the apex, slightly ciliate. Corolla yellow or greenish-yellow, blotched with 5 dark brownish violet or purplish markings strongly contrasting with the surrounding limb, sometimes spreading downwards and somewhat along the nerves upwards, 4–7(8) mm long, rotate; tube subglabrous, furnished with few ± sessile glands, on the inside with felted indumentum near the insertion of the stamens and with a ring of hyaline hairs at the mouth; limb (5)6–9 mm across, shortly 5-lobed, erect to widely spreading, sometimes reflexed when fully expanded, on the outside with ± appressed, short hairs and also sessile glands often along the nerves, on the median region of each lobe on the part not folded in bud, glabrous inside except for the lobes with minute hairs towards the apex, slightly ciliate
Morphology Reproductive morphology Flowers Gynoecium Pistil
Ovary 1.2–2 × 0.8–1.5 mm, ovoid or subglobose, glabrous; style greenish-blue, (1)1.5–2.5(3) mm long, filiform, straight or curved upwards.
Morphology Reproductive morphology Seeds
Seeds whitish to yellowish, 1.7–2.2 × 1.5–1.7 mm, elliptic or ± orbicular in outline, sometimes reniform, reticulate-foveolate. Seeds whitish to yellowish, 1. 7–2.2 × 1. 5–1. 7 mm, elliptic or ± orbicular in outline, sometimes reniform, reticulate-foveolate
Cytology
Chromosome number: 2n=24.

Native to:

Mexico Northeast, Mexico Northwest, Mexico Southwest

Introduced into:

Botswana, Egypt, Eritrea, Ethiopia, Greece, India, New Mexico, New South Wales, Norfolk Is., Portugal, Primorye, Queensland, Spain, Uzbekistan, West Siberia, Zambia, Zaïre, Zimbabwe

Physalis ixocarpa Brot. ex Hornem. appears in other Kew resources:

First published in Hort. Bot. Hafn., Suppl.: 26 (1819)

Accepted by

  • Allred, K.W. (2012). Flora Neomexicana, ed. 2, 1: 1-599. Range Science Herbarium, Las Cruces, New Mexico.
  • Barkalov, V.Y. (ed.) (1991). Plantae Vasculares Orientalis Extremi Sovietici 5: 1-388. Nauka, Leningrad.
  • Boulos, L. (2002). Flora of Egypt 3: 1-373. Al Hadara Publishing, Cairo.
  • Castroviejo, S. (ed.) in Castroviejo, S. & al. (eds.) (2012). Flora Iberica 11: 1-672. Real Jardín Botánico, CSIC, Madrid.
  • Dimopoulos, P., Raus, T., Bergmeier, E., Constantinidis, T., Iatrou, G., Kokkini, S., Strid, A., & Tzanoudakis, D. (2013). Vascular plants of Greece. An annotated checklist: 1-372. Botanic gardens and botanical museum Berlin-Dahlem, Berlin and Hellenic botanical society, Athens.
  • Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • Orchard, A.E. (ed.) (1994). Oceanic Islands 1 Flora of Australia 49: 1-681. Australian Government Publishing Service, Canberra.
  • Pretz, C. & Deanna, R. (2020). Typifications and nomenclatural notes on Physalis (Solanaceae) from the United States Taxon 69: 170-192.
  • Purdie, R.W., Symon, D.E. & Haegi, L. (1982). Flora of Australia 29: 1-208. Australian Government Publishing Service, Canberra.
  • Sheremetova, S.A., Ebel, A.L. & Buko, T.E. (2011). Supplement to the flora of Kemerovo region since 2001 till 2010 Turczaninowia 14(1): 65-74.
  • Vvedensky, A.I. (ed.) (1961). Flora Uzbekistana 5: 1-667. Izd-va Akademii nauk Uzbekskoi SSR, Tashkent.

Not accepted by

  • Dobignard, A. & Chatelain, C. (2013). Index synonymique de la flore d'Afrique du nord 5: 1-451. Éditions des conservatoire et jardin botaniques, Genève. [Cited as Physalis philadelphica.]

Literature

Flora Zambesiaca

  • Agnew, Upland Kenya Wild Fl., ed. 2: 244 (1994).
  • Bol. Soc. Brot., Sér. 2, 44: 344, 357, t. 2, t. 4–8, t. 10 fig. A–L (1970).
  • Garcia de Orta 17: 286 quadro I (1969)
  • ex A. DC., Cat. Pl. Hort. Bot. Monsp.: 50 (1813) as “ixocarpos” nom. nud.
  • ex Hornem., Hort. Bot. Hafn., Suppl.: 26 (1819).

Kew Backbone Distributions

  • Allred, K.W. (2012). Flora Neomexicana, ed. 2, 1: 1-599. Range Science Herbarium, Las Cruces, New Mexico.
  • Barkalov, V.Y. (ed.) (1991). Plantae Vasculares Orientalis Extremi Sovietici 5: 1-388. Nauka, Leningrad.
  • Boulos, L. (2002). Flora of Egypt 3: 1-373. Al Hadara Publishing, Cairo.
  • Castroviejo, S. (ed.) in Castroviejo, S. & al. (eds.) (2012). Flora Iberica 11: 1-672. Real Jardín Botánico, CSIC, Madrid.
  • Dimopoulos, P., Raus, T., Bergmeier, E., Constantinidis, T., Iatrou, G., Kokkini, S., Strid, A., & Tzanoudakis, D. (2013). Vascular plants of Greece. An annotated checklist: 1-372. Botanic gardens and botanical museum Berlin-Dahlem, Berlin and Hellenic botanical society, Athens.
  • Gonçalves, A.E. (2005). Flora Zambesiaca 8(4): 1-124. Royal Botanic Gardens, Kew.
  • Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • Orchard, A.E. (ed.) (1994). Oceanic Islands 1 Flora of Australia 49: 1-681. Australian Government Publishing Service, Canberra.
  • Purdie, R.W., Symon, D.E. & Haegi, L. (1982). Flora of Australia 29: 1-208. Australian Government Publishing Service, Canberra.
  • Sheremetova, S.A., Ebel, A.L. & Buko, T.E. (2011). Supplement to the flora of Kemerovo region since 2001 till 2010 Turczaninowia 14(1): 65-74.
  • Vvedensky, A.I. (ed.) (1961). Flora Uzbekistana 5: 1-667. Izd-va Akademii nauk Uzbekskoi SSR, Tashkent.

Flora Zambesiaca
Flora Zambesiaca
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Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

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