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This species is accepted, and its native range is Mexico to Venezuela and Argentina (Jujuy), Cuba.
Physalis lagascae

[FZ]

Flora Zambesiaca. Vol. 8, Part 4. Solanaceae. Gonçalves AE. 2005

Morphology General
Erect to decumbent or prostrate, weak or occasionally ± robust, ± dichotomously much branched, annual herb, (0.05)0.1–1 m high (elsewhere said to reach 1.5 m), often tinged purple, ± clothed all over with simple, stout, often multicellular, white or hyaline, ± spreading, usually eglandular hairs 1–4 mm long, and also rigid, curved at the tip, appressed to spreading, often minute, hairs, furnished with ± sessile glands too
Morphology Reproductive morphology Flowers Pedicel
Pedicel 2–5(9) mm long, villous or puberulous, occasionally subglabrous, in fruit elongated to 9(10) mm
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary 0.8–0.9 × 0.6–0.7 mm, ellipsoid or ± globose, glabrous.
Morphology Reproductive morphology Flowers Gynoecium Style
Style 2–3 mm long, filiform, ± curved upwards
Note
The specimens Torre et al. in Torre 17705, 18035 and 19042, from the Tete Province of Mozambique have relatively large leaves, with petioles up to 8 cm long and laminas up to 10.5 × 7.5 cm. The specimen Flanagan 3305, from Zimbabwe, referred to by Wild, loc. cit. (1953) as P. minima, but not seen by me, is probably this species. Chromosome number: 2n=24
Distribution
Mozambique Zambia Zimbabwe Malawi ZAM N, ZAM W, ZAM C, ZAM S, ZIM N, ZIM E, MAL C, MAL S, MOZ Z, MOZ T, MOZ MS Native to tropical America (Mexico and Antilles), now extending northwards to the United States and southwards throughout Central America and the Caribbean to Bolivia.
Morphology General Habit
Erect to decumbent or prostrate, weak or occasionally ± robust, ± dichotomously much branched, annual herb, (0.05)0.1–1 m high (elsewhere said to reach 1.5 m), often tinged purple, ± clothed all over with simple, stout, often multicellular, white or hyaline, ± spreading, usually eglandular hairs 1–4 mm long, and also rigid, curved at the tip, appressed to spreading, often minute, hairs, furnished with ± sessile glands too.
Morphology Branches
Branches angular or angular-ribbed, striate, drying ± sulcate, sparingly villous, with ± long hairs mainly on the emergent parts and also short and minute hairs somewhat localized near the nodes, occasionally subglabrous. Branches angular or angular-ribbed, striate, drying ± sulcate, sparingly villous, with ± long hairs mainly on the emergent parts and also short and minute hairs somewhat localized near the nodes, occasionally subglabrous
Morphology Leaves
Leaves solitary or geminate; petiole 0.5–4.5(8) cm long, often slightly winged, rather sheathing at the base; lamina membranous or ± fleshy, 1.5–7.5(10.5) × 0.8–4.5(7.5) cm, ovate to lanceolate, sometimes rhombic, ovate-oblong, elliptic or oblanceolate, base shortly cuneate or attenuate, sometimes obtuse, rounded, truncate or subcordate, and often oblique or unequal-sided, sometimes ± decurrent into the petiole, apex acuminate, acute or obtuse, entire to somewhat coarsely sinuate-dentate, rarely dentate, the teeth unequal, ± triangular, obtuse, the sinuses rounded, both surfaces ± sparsely pilose or subglabrous but pubescent along the nerves and near the margins; minor leaves with lamina 0.5–0.7 × 0.3–0.4 cm, elliptic, sometimes present. Leaves solitary or geminate; petiole 0.5–4.5(8) cm long, often slightly winged, rather sheathing at the base; lamina membranous or ± fleshy, 1. 5–7.5(10.5) × 0.8–4.5(7.5) cm, ovate to lanceolate, sometimes rhombic, ovate-oblong, elliptic or oblanceolate, base shortly cuneate or attenuate, sometimes obtuse, rounded, truncate or subcordate, and often oblique or unequal-sided, sometimes ± decurrent into the petiole, apex acuminate, acute or obtuse, entire to somewhat coarsely sinuate-dentate, rarely dentate, the teeth unequal, ± triangular, obtuse, the sinuses rounded, both surfaces ± sparsely pilose or subglabrous but pubescent along the nerves and near the margins; minor leaves with lamina 0.5–0.7 × 0.3–0.4 cm, elliptic, sometimes present
Morphology Reproductive morphology Flowers
Flowers solitary, inserted by the side of the petiole-base appearing axillary, erect to pendulous; pedicel 2–5(9) mm long, villous or puberulous, occasionally subglabrous, in fruit elongated to 9(10) mm. Flowers solitary, inserted by the side of the petiole-base appearing axillary, erect to pendulous.
Morphology Reproductive morphology Flowers Calyx
Calyx (1.5)2–3.5(4) mm long, 1.5–2.5(3) mm across at the base of the lobes, campanulate, sub-angled-ribbed, truncate or invaginated at the base, with scattered, stout, white or hyaline, ± spreading, long hairs, or puberulous especially along the ribs, on the inside glabrous except for the lobes with ± dense, minute indumentum towards the apex, near the margins and sometimes also along the midribs; lobes subequal, (0.3)0.5–1.2(1.5) × 0.7–1(1.2) mm, ± deltate or ovate-triangular to triangular-lanceolate, acute or obtuse, sometimes sub-acuminate, rarely sub-truncate; in fruit greenish, usually drying purple-veined, (10)12–20(23) × (8)10–15(20) mm, globose to ± ovoid, 10-angled-ribbed or 10-ribbed, slightly obtuse or apiculate at the summit, half to almost filled by the fruit, subglabrous to pilose, mostly with long hairs especially along the ribs and nerves, often the ribs with widely spaced small teeth or enations derived from the hair bases, the lobes (0.5)1–2.5(3) × 1–2 mm. Calyx (1. 5)2–3.5(4) mm long, 1. 5–2.5(3) mm across at the base of the lobes, campanulate, sub-angled-ribbed, truncate or invaginated at the base, with scattered, stout, white or hyaline, ± spreading, long hairs, or puberulous especially along the ribs, on the inside glabrous except for the lobes with ± dense, minute indumentum towards the apex, near the margins and sometimes also along the midribs; lobes subequal, (0.3)0.5–1. 2(1. 5) × 0.7–1(1. 2) mm, ± deltate or ovate-triangular to triangular-lanceolate, acute or obtuse, sometimes sub-acuminate, rarely sub-truncate; in fruit greenish, usually drying purple-veined, (10)12–20(23) × (8)10–15(20) mm, globose to ± ovoid, 10-angled-ribbed or 10-ribbed, slightly obtuse or apiculate at the summit, half to almost filled by the fruit, subglabrous to pilose, mostly with long hairs especially along the ribs and nerves, often the ribs with widely spaced small teeth or enations derived from the hair bases, the lobes (0.5)1–2.5(3) × 1–2 mm
Morphology Reproductive morphology Flowers Corolla
Corolla greenish-yellow or yellow to white or greenish-cream, slightly purplish marked or apparently unmarked, more rarely blotched with 5 ± dark purple to brown markings not strongly contrasting with the surrounding limb, (3)4–5(7) mm long, tubular-campanulate, often narrowly so; tube glabrous, on the inside with a few hyaline hairs from the insertion of the stamens upwards; limb (2)3–6 mm across, shortly 5-lobed, erect or spreading, rarely reflexed when fully expanded, glabrous or puberulous on the parts not folded in bud, glabrous inside, ciliate. Corolla greenish-yellow or yellow to white or greenish-cream, slightly purplish marked or apparently unmarked, more rarely blotched with 5 ± dark purple to brown markings not strongly contrasting with the surrounding limb, (3)4–5(7) mm long, tubular-campanulate, often narrowly so; tube glabrous, on the inside with a few hyaline hairs from the insertion of the stamens upwards; limb (2)3–6 mm across, shortly 5-lobed, erect or spreading, rarely reflexed when fully expanded, glabrous or puberulous on the parts not folded in bud, glabrous inside, ciliate
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens included or slightly exserted, equal or subequal; filaments (1)1.5–3(3.5) mm long, filiform, attached to the corolla tube near the base, furnished with few hairs; anthers yellowish, sometimes blue-margined, 0.8–1 mm long, ± oblong or elliptic in outline, straight after anthesis. Stamens included or slightly exserted, equal or subequal; filaments (1)1. 5–3(3.5) mm long, filiform, attached to the corolla tube near the base, furnished with few hairs; anthers yellowish, sometimes blue-margined, 0.8–1 mm long, ± oblong or elliptic in outline, straight after anthesis
Morphology Reproductive morphology Flowers Disc
Disk 0.2–0.3 mm high, fleshy, glabrous. Disk 0.2–0.3 mm high, fleshy, glabrous
Morphology Reproductive morphology Flowers Gynoecium Pistil
Ovary 0.8–0.9 × 0.6–0.7 mm, ellipsoid or ± globose, glabrous; style 2–3 mm long, filiform, ± curved upwards.
Morphology Reproductive morphology Fruits
Fruit greenish-yellow or pale yellow, occasionally yellow, subsessile or with a gynobase up to 1 mm long on the invaginated base of the erect to pendulous calyx, (5)6–10 mm in diameter, ± globose or slightly ovoid, viscid. Fruit greenish-yellow or pale yellow, occasionally yellow, subsessile or with a gynobase up to 1 mm long on the invaginated base of the erect to pendulous calyx, (5)6–10 mm in diameter, ± globose or slightly ovoid, viscid
Morphology Reproductive morphology Seeds
Seeds light brown or yellowish, (1.5)1.8–2 × (1.5)1.6–1.8(2) mm, ovate to orbicular in outline, sometimes reniform, reticulate-foveolate. Seeds light brown or yellowish, (1. 5)1. 8–2 × (1. 5)1. 6–1. 8(2) mm, ovate to orbicular in outline, sometimes reniform, reticulate-foveolate
Cytology
Chromosome number: 2n=24.

[FTEA]

Solanaceae, Jennifer M Edmonds. Oliganthes, Melongena & Monodolichopus, Maria S. Vorontsova & Sandra Knapp. Flora of Tropical East Africa. 2012

Type
Type: based on P. parviflora sensu Lagasca, Gen. Sp. Nov. Diagn.: 11, n. 147 (1816); type not cited
Morphology General Habit
Annual herb usually < 0.5 m high, occasionally taller, erect, decumbent, prostrate or creeping.
Morphology Stem
Main stems sparsely to densely branched, green or purple-tinged, pilose with simple hairs (in our area)
Morphology Leaves
Leaves light to dark green, 1.7–4.5(–6) × 1–3(–4.2) cm, smaller above, ovate to ovate-lanceolate, bases cuneate, often oblique, sometimes decurrent, margins entire to sinuate, rarely sinuate-dentate, apices acute, sparsely to moderately pilose (in our area), hairs usually short and appressed, denser on veins, midribs and lower surfaces; petioles 0.5–3.2 cm, pubescence as stems
Morphology Reproductive morphology Flowers
Flowers solitary, small, in leaf or branch axils; pedicels 1.5–4 mm long, erect in flower, pubescent, 3–7 mm long and erect to recurved in fruit. Calyx cupulate, 1.7–3(–4) mm long with five broadly triangular acute lobes 0.5–1 × 0.5–1.5 mm with ciliate margins, often with long spreading white hairs, enlarged and persistent in fruit with connivent lobes 0.8–1.8 × 0.8–2 mm. Corolla white, cream, pale yellow or green, tubular/campanulate, 2.9–5 × 3–4 mm diameter, shortly pubescent externally, margin ciliate, undulating or with five broadly triangular obtuse lobes 0.2–1 × 0.7–1.1 mm
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens often visible in corolla throat; filaments free for 1.5–2.5(–3) mm; anthers pale yellow, ovoid to oblong, bilobed, 0.5–1.3 × 0.3–0.5 mm
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary brownish, ovoid, 0.7–1.1 × 0.5–1.1 mm, smooth, glabrous; disc 0.7–1.5 mm diameter; style often exserted, 2–2.5 mm long; stigma 0.2–0.3 × 0.2–0.5 mm
Morphology Reproductive morphology Fruits
Fruit pale green to yellow, globose, rarely ovoid, 5–10 mm diameter, smooth, subsessile or on inverted gynobase (–0.75 mm) of enlarged and inflated reticulately-veined pale green globose calyx, 0.8–1.8 × 0.7–2 cm, the mouth closed by connivent calyx lobes, veins often clothed with long spreading hairs
Morphology Reproductive morphology Seeds
Seeds yellow, orbicular, 1.2–1.8 × 1.1–1.7 mm, compressed
Figures
Fig 15/1–4, p 72
Ecology
Weed of cultivation, plantations and abandoned areas, also on river-banks, disturbed sites, edges of paddy-fields, grass patches in floodplains; common to local; 40–1850 m
Conservation
Widespread; least concern (LC)
Note
This taxon is widely known as P. minima L. – an appropriate epithet in view of its characteristic small leaves, flowers, berries and fruiting calyces. However, typification of this Linnaean species is extremely complex with the lectotype of P. minima (Hort. Cliff. n. 62, Physalis 5 (BM!)) being conspecific with P. angulata, with which it has been synonymised. Moreover, most of the descriptions and plates on which Linnaeus based this species (e.g. Herm. Lugdb.: 569, t. 571; and Rheede Mal. 10, t. 140, f. 71) also refer to P. angulata rather than the small-flowered prostrate plant described here. Numerous authors have discussed the difficulties of correctly naming and typifying this taxon ( cf. Fernandes in Garcia de Orta, 17: 282 (1969)), and some of the synonyms cited in the literature for P. minima belong to P. angulata. Though the name P. lagascae has been used here in line with various recent generic treatments, there are no known type specimens and the protologue does not entirely reflect the morphology characterised by East African specimens. Roemer & Schultes (1819) described the branches of their taxon as villous, repeating the phrase used by Opiz (1817) when describing P. parviflora on which P. lagascae was based. While the majority of East African specimens are sparsely to moderately pilose, there are a number of specimens from Kenya and Tanzania which are densely villous even to the naked eye. In other features these specimens are very similar to the species described above and they have been included under P. lagascae. Indeed, hairy and glabrescent varieties of this species have been recognised as var. lagascae and var. glabrescens O.E. Schulz respectively in Central America ( cf. Schulz, 1909; Waterfall, 1967). These villous specimens could, however, belong to the taxon described as P. pubescens L. in Flora Zambesiaca (apparently based on a single specimen from Zambia). It is possible that following more detailed work on its pubescence variation, a name change might become necessary for P. lagascae. The fruits of this species are edible, and the boiled leaves are eaten as a vegetable in U 1, K 2 and Ethiopia; it is considered to be a medicinal plant throughout its range.
Distribution
Flora districts: U1 U2 U3 U4 K2 K4 K5 K ?7; T1 T2 T3 T4 T ?5, T6 T7 Range: Probably native to Mexico but now a cosmopolitan weed throughout tropical and subtropical regions

[CPLC]

Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

Distribution
Nativa en Colombia; Alt. 100 - 438 m.; Valle del Magdalena.
Morphology General Habit
Hierba, subarbusto
Conservation
No Evaluada

Native to:

Argentina Northwest, Bolivia, Colombia, Costa Rica, Cuba, Ecuador, El Salvador, Guatemala, Honduras, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Nicaragua, Panamá, Peru, Trinidad-Tobago, Venezuela

Introduced into:

Andaman Is., Angola, Assam, Bangladesh, Benin, Bismarck Archipelago, Botswana, Burkina, Burundi, Cambodia, Cameroon, Cape Verde, Caroline Is., Central African Repu, Chad, China South-Central, China Southeast, Christmas I., Cocos (Keeling) Is., Congo, Equatorial Guinea, Eritrea, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., Gulf States, India, Ivory Coast, Jawa, Kenya, Laccadive Is., Laos, Liberia, Libya, Madagascar, Malawi, Malaya, Maldives, Mali, Marianas, Mauritania, Mauritius, Mozambique, Myanmar, Nauru, New Caledonia, New Guinea, Nicobar Is., Niger, Nigeria, Niue, Oman, Philippines, Rwanda, Samoa, Senegal, Sierra Leone, Socotra, Solomon Is., Sri Lanka, Sudan, Tanzania, Thailand, Togo, Tonga, Tubuai Is., Uganda, Vietnam, Wallis-Futuna Is., West Himalaya, Yemen, Zambia, Zaïre, Zimbabwe

Physalis lagascae Roem. & Schult. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Feb 1, 2002 Cheek, M. [8906], Cameroon K000109667
Pringle, C.G. [11055], Mexico K000042260
Hinton, G.B. [8493], Mexico K000042265
Hinton, G.B. [2717], Mexico K000042259
Pringle, C.G. [8447], Mexico K000042258
Hinton, G.B. [5204], Mexico K000042262
Hinton, G.B. [8418], Mexico K000042264
Langlassé, E. [393], Mexico K000042263
Hinton, G.B. [5982], Mexico K000042266
Hinton, G.B. [5821], Mexico K000042261

First published in Syst. Veg., ed. 15 bis 4: 679 (1819)

Accepted by

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Literature

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