Trema orientale (L.) Blume

Native to:

Andaman Is., Angola, Assam, Bangladesh, Benin, Bismarck Archipelago, Borneo, Burkina, Burundi, Cambodia, Cameroon, Cape Provinces, Caroline Is., Central African Republic, Chad, China South-Central, China Southeast, Comoros, Congo, Cook Is., Djibouti, DR Congo, East Himalaya, Eritrea, Eswatini, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., Hainan, India, Ivory Coast, Japan, Jawa, Kazan-retto, Kenya, KwaZulu-Natal, Laos, Lesser Sunda Is., Liberia, Madagascar, Malawi, Malaya, Mali, Maluku, Marianas, Marquesas, Mozambique, Myanmar, Namibia, Nansei-shoto, Nepal, New Guinea, Nicobar Is., Nigeria, Niue, Northern Provinces, Northern Territory, Ogasawara-shoto, Oman, Philippines, Queensland, Rwanda, Samoa, Santa Cruz Is., Saudi Arabia, Senegal, Seychelles, Sierra Leone, Solomon Is., Somalia, South China Sea, Sri Lanka, Sudan-South Sudan, Sulawesi, Sumatera, Tanzania, Thailand, Tibet, Togo, Tonga, Uganda, Vietnam, West Himalaya, Yemen, Zambia, Zimbabwe

Introduced into:

Germany, Hawaii, Mauritius, Réunion

POWO follows these authorities in accepting this name:

• Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
• Banks, R.C. (ed.) (1982). Wildlife and wildlife habitat of American Samoa 2: 1-151. U.S. Fisch and Wildlife services, Washington.
• Barooah, C. & Ahmed, I. (2014). Plant diversity of Assam. A checklist of Angiosperms and Gymnosperms: 1-599. Assam science technology and environment council, India.
• Bosser, J., Cadet, T., Guého, J. & Marais, W. (eds.) (1985). Flore des Mascareignes 161-169: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
• Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
• Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 1: 1-114. ORSTROM, Bangui.
• Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 3: 1-89. ORSTROM, Bangui.
• Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
• Caminho, J., Mutemba, L., Matusse, E., Osborne, J., Wursten, B., Burrows, J., Cianciullo, S., Malatesta, L. & Attorre, F. (2022). An updated checklist of Mozambique’s vascular plants. PhytoKeys 189: 61-80.
• Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau). Garcia de Orta, Série de Botânica 17: 97-141.
• Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
• Choudhary, R.K., Srivastava, R.C., Das, A.K. & Lee, J. (2012). Floristic diversity assessment and vegetation analysis of Upper Siang district of eastern Himalaya in North East India. Korean Journal of Plant Taxonomy 42: 222-246.
• Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
• Dassanayake (ed.) (1999). A Revised Handbook to the Flora of Ceylon 13: 1-284. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
• Figueiredo, E. & Smith, G.F. (2008). Plants of Angola. Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
• Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya. Journal of East African Natural History 99: 129-226.
• George, A.S. (ed.) (1989). Flora of Australia 3: 1-219. Australian Government Publishing Service, Canberra.
• Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist. Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
• Ghazanfar, S.A. (1992). An Annotated Catalogue of the Vascular Plants of Oman and their Vernacular names. Scripta Botanica Belgica 2: 1-153.
• Gosline, G., Bidault, E., van der Burgt, X., Cahen, D., Challen, G., Condé, N., Couch, C., Couvreur, T.L.P., Dagallier, L.M.J., Darbyshire, I., Dawson, S., Doré, T.S., Goyder, D., Grall, A., Haba, P., Haba, P., Harris, D., Hind, D.J.N., Jongkind, & al. (2023). A Taxonomically-verified and Vouchered Checklist of the Vascular Plants of the Republic of Guinea. Nature, scientific data 10, Article number: 327: [1]-[12].
• Grierson, A.J.C. & Long, D.G. (1983). Flora of Bhutan 1(1): 1-186. Royal Botanic Gardens, Edinburgh.
• Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands. Research Bulletin Dodo Creek Research Station 7: 1-203.
• Harris, D.J. (2002). Vasc. Pl. Dzanga-Sangha: 1-274. National Botanic Garden (Belgium), Meise.
• Hassler, M. & Muer, T. (2022). Flora Germanica: alle Farn- und Blütenpflanzen Deutschlands in Text und Bild 1: 1-864. Verlag Regionalkultur, Uberstadt-Weiher.
• Hedberg, I. & Edwards, S. (eds.) (1989 publ. 1990). Flora of Ethiopia and Eritrea 3: 1-659. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
• Iwatsuki, K., Boufford, D.E. & Ohba, H. (eds.) (2006). Flora of Japan IIa: 1-550. Kodansha Ltd., Tokyo.
• Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
• Kobayashi, S. & Ono, M. (1987). A Revised List of Vascular Plants Indigenous and Introduced to the Bonin (Ogasawara) and the Volcano (Kazan) Islands. Ogasawara Research 13: 1-55.
• Kotiya, A., Solanki, Y. & Reddy, G.V. (2020). Flora of Rajasthan: 1-769. Rajasthan state biodiversity board.
• Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar. Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
• Launert, E. & Cope, G.V. (eds.) (1991). Flora Zambesiaca 9(6): 1-135. Royal Botanic Gardens, Kew.
• Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
• Lebrun, J.P., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
• Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo). Taxonomania 30: 1-307.
• Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
• Lorence, D.H. & Wagnwe, W.L. (2020). Flora of the Marquesas Islands 2: 413-1135. National Tropical Botanic Garden, Smithsonian, DRPF.
• Lê, T.C. (2003). Danh lục các loài thực vật Việt Nam 2: 1-1203. Hà Nội : Nhà xuất bản Nông nghiệp.
• Mannheimer, C.A. & Curtis, B.A. (eds.) (2009). Le Roux and Müller's field guide to the trees and shrubs of Namibia, rev. ed.: 1-525. Macmillan Education Namibia, Windhoek.
• Masharabu, T., Bigendako, M.J., Lejoly, J., Nkengurutse, J., Noret, N., Bizuru, E. & Bogaert, J. (2010). Etude analytique de la flore et de la végétation du Parc National de la Ruvubu, Burundi. International Journal of Biological and Chemical Sciences 4: 834-856.
• Miller, A.G. & Cope, T.A. (1996). Flora of the Arabian peninsula and Socotra 1: 1-586. Edinburgh university press.
• Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. & Armstrong, K. (2007). A checklist of the vascular plants of Lao PDR: 1-394. Royal Botanic Gardens, Edinburgh.
• Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India. Journal of Economic and Taxonomic Botany 32: 403-500.
• Pasha, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
• Polhill, R.M. (1966). Flora of Tropical East Africa, Ulmaceae: 1-14.
• Pusalkar, P.K., Mao, A.A. & Inge, P. (eds.) (2022). Flora of India 24: 1-688. Botanical Survey of India.
• Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon. Scripta Botanica Belgica 35: 1-438.
• Sosef, M.S.M. & al. (eds.) (2022). Flore du Gabon 58: 1-150. Muséum National D'Histoire Naturelle, Paris; Margraf Publishers, Weikersheim; Meise Botanic Garden.
• Sykes, W.R. (2016). Flora of the Cook Islands: 1-973. National Tropical Botanical Garden, Hawaii.
• Thulin, M. (ed.) in Thulin, M. (ed.) (1999). Flora of Somalia 2: 1-303. The Royal Botanic Gardens, Kew.
• Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.
• Toyama, H. & al. (2013). Inventory of the woody flora in Permanent plats of Kampong Thom and Kompong Chhnang provinces, Cambodia. Acta Phytotaxonomica et Geobotanica 64: 45-105.
• Troupin, G. (ed.) (1978). Flora du Rwanda 1: 1-413. Musee Royal de l'Afrique Centrale.
• Turner, I.M. (1995 publ. 1997). A catalogue of the Vascular Plants of Malaya. Gardens' Bulletin Singapore 47(2): 347-655.
• Velayos, M., Barberá, P., Cabezas, F.J., de la Estrella, M., Fero, M. & Aedo, C. (2014). Checklist of the vascular plants of Annobón (Equatorial Guinea). Phytotaxa 171: 1-78.
• Wiriadinata, H., Girmansyah, D., Hunter, J.M., Hoover, W.S. & Kartawinata, K. (2013). Floristic study of West Sumbawa, Indonesia. Reinwardtia 13: 391-404.
• Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
• van Steenis, C.G.G.J. (ed.) (1974-1978). Flora Malesiana 8: 1-577. Noordhoff-Kolff N.V., Djakarta.

Kew Backbone Distributions

• Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
• Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
• Balkrishna, A. (2018). Flora of Morni Hills (Research & Possibilities): 1-581. Divya Yoga Mandir Trust.
• Banks, R.C. (ed.) (1982). Wildlife and wildlife habitat of American Samoa 2: 1-151. U.S. Fisch and Wildlife services, Washington.
• Barooah, C. & Ahmed, I. (2014). Plant diversity of Assam. A checklist of Angiosperms and Gymnosperms: 1-599. Assam science technology and environment council, India.
• Bosser, J., Cadet, T., Guého, J. & Marais, W. (eds.) (1985). Flore des Mascareignes 161-169: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
• Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
• Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 1: 1-114. ORSTROM, Bangui.
• Castle, G.E. (1994). Flore des Seychelles Dicotylédones: 1-663. Orstom Editions.
• Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau). Garcia de Orta, Série de Botânica 17: 97-141.
• Choudhary, R.K., Srivastava, R.C., Das, A.K. & Lee, J. (2012). Floristic diversity assessment and vegetation analysis of Upper Siang district of eastern Himalaya in North East India. Korean Journal of Plant Taxonomy 42: 222-246.
• Collenette, S. (1999). Wildflowers of Saudi Arabia: 1-799. National commission for wildlife conservation and development (NCWCD), Kingdom of Saudi Arabia.
• Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
• Dassanayake (ed.) (1999). A Revised Handbook to the Flora of Ceylon 13: 1-284. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
• Figueiredo, E. & Smith, G.F. (2008). Plants of Angola. Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
• Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya. Journal of East African Natural History 99: 129-226.
• Fosberg, F.R., Sachet, M.-H., Oliver, R. (1979). A geographical checklist of the Micronesian Dicotyledonae. Micronesica; Journal of the College of Guam 15: 41-295.
• George, A.S. (ed.) (1989). Flora of Australia 3: 1-219. Australian Government Publishing Service, Canberra.
• Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist. Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
• Ghazanfar, S.A. (1992). An Annotated Catalogue of the Vascular Plants of Oman and their Vernacular names. Scripta Botanica Belgica 2: 1-153.
• Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands. Research Bulletin Dodo Creek Research Station 7: 1-203.
• Hassler, M. & Muer, T. (2022). Flora Germanica: alle Farn- und Blütenpflanzen Deutschlands in Text und Bild 1: 1-864. Verlag Regionalkultur, Uberstadt-Weiher.
• Hedberg, I. & Edwards, S. (eds.) (1989 publ. 1990). Flora of Ethiopia and Eritrea 3: 1-659. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
• Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
• Iwatsuki, K., Boufford, D.E. & Ohba, H. (eds.) (2006). Flora of Japan IIa: 1-550. Kodansha Ltd., Tokyo.
• Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
• Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
• Kobayashi, S. & Ono, M. (1987). A Revised List of Vascular Plants Indigenous and Introduced to the Bonin (Ogasawara) and the Volcano (Kazan) Islands. Ogasawara Research 13: 1-55.
• Launert, E. & Cope, G.V. (eds.) (1991). Flora Zambesiaca 9(6): 1-135. Royal Botanic Gardens, Kew.
• Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
• Lebrun, J.P., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
• Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo). Taxonomania 30: 1-307.
• Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
• Lorence, D.H. & Wagnwe, W.L. (2020). Flora of the Marquesas Islands 2: 413-1135. National Tropical Botanic Garden, Smithsonian, DRPF.
• Mannheimer, C.A. & Curtis, B.A. (eds.) (2009). Le Roux and Müller's field guide to the trees and shrubs of Namibia, rev. ed.: 1-525. Macmillan Education Namibia, Windhoek.
• Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India. Journal of Economic and Taxonomic Botany 32: 403-500.
• Pasha, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
• Polhill, R.M. (1966). Flora of Tropical East Africa, Ulmaceae: 1-14.
• Pusalkar, P.K., Mao, A.A. & Inge, P. (eds.) (2022). Flora of India 24: 1-688. Botanical Survey of India.
• Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
• Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon. Scripta Botanica Belgica 35: 1-438.
• Sykes, W.R. (1970). Contributions to the flora of Niue. Bulletin, New Zealand Department of Scientific and Industrial Research 200: 1-321.
• Sykes, W.R. (2016). Flora of the Cook Islands: 1-973. National Tropical Botanical Garden, Hawaii.
• Thulin, M. (ed.) in Thulin, M. (ed.) (1999). Flora of Somalia 2: 1-303. The Royal Botanic Gardens, Kew.
• Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.
• Toyama, H. & al. (2013). Inventory of the woody flora in Permanent plats of Kampong Thom and Kompong Chhnang provinces, Cambodia. Acta Phytotaxonomica et Geobotanica 64: 45-105.
• Troupin, G. (ed.) (1978). Flora du Rwanda 1: 1-413. Musee Royal de l'Afrique Centrale.
• Turner, I.M. (1995 publ. 1997). A catalogue of the Vascular Plants of Malaya. Gardens' Bulletin Singapore 47(2): 347-655.
• Wagner, W.L., Herbst, D.R. & Sohmer, S.H. (1999). Manual of the Flowering Plants of Hawai'i, rev. ed., 2: 989-1918. University of Hawai'i Press, Bishop Museum Press.
• Wiriadinata, H., Girmansyah, D., Hunter, J.M., Hoover, W.S. & Kartawinata, K. (2013). Floristic study of West Sumbawa, Indonesia. Reinwardtia 13: 391-404.
• Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
• Wu, Z. & Raven, P.H. (eds.) (2003). Flora of China 5: 1-505. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
• van Steenis, C.G.G.J. (ed.) (1974-1978). Flora Malesiana 8: 1-577. Noordhoff-Kolff N.V., Djakarta.

Flora of Somalia

• Flora Somalia, Vol 2, (1999) Author: by S. Fici [updated by M. Thulin 2008]

Flora of Tropical East Africa

• Bl., Mus. Bot. Lugd.-Bat. 2: 62 (1852).
• Leroy in Fl. Madag. 54: 10 (1952).
• Polhill in Kew Bulletin 19: 143 (1964).