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This species is accepted, and its native range is Tanzania to Zimbabwe.
A specimen from Kew's Herbarium

[FZ]

Flora Zambesiaca. Vol 8, Pt 7. Avicenniaceae, R. Fernandes. Nesogenaceae, M.A. Diniz. Verbenaceae, R. Fernandes. Lamiaceae, R. Fernandes. 2005.

Type
Type from Tanzania.
Morphology General Habit
Tree c. 15–18 m tall; bark white or pale brown, smooth; older branches dark red-brown; young branches crisply pilose Tree c. 15–18 m tall; bark white or pale brown, smooth; older branches dark red-brown; young branches crisply pilose.
Morphology Leaves
Leaves drying dark, 3–5-foliolate; petioles 9–15 cm long, terete; median petiolules 0.5–2.5 cm long; leaflets 10–20 × 5–6.5 cm, median leaflet largest, elliptic to broad obovate, rounded or apiculate to shortly acuminate at the apex, cuneate to ± attenuate at the base, entire to remotely serrulate, glabrous on both surfaces with a few short hairs on the nerves beneath, nerves ± raised on both surfaces Leaves drying dark, 3–5-foliolate; petioles 9–15 cm long, terete; median petiolules 0.5–2.5 cm long; leaflets 10–20 × 5–6.5 cm, median leaflet largest, elliptic to broad obovate, rounded or apiculate to shortly acuminate at the apex, cuneate to ± attenuate at the base, entire to remotely serrulate, glabrous on both surfaces with a few short hairs on the nerves beneath, nerves ± raised on both surfaces.
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary obovoid, sparsely gland-dotted at apex only, glabrous, truncate at apex; style 5 mm long
Ecology
In the Flora Zambesiaca area known only from the evergreen rain forest at Chirinda in Zimbabwe; c. 1100 m.
Note
Only four gatherings of this species from the Flora Zambesiaca region have been seen by this author – all from the Chirinda Forest in Zimbabwe.  This rain forest species was apparently first collected there in 1948 (by N.C. Chase 435, Jan 1948, Chirinda Forest) and wrongly determined as V. lokundjensis W. Piep. by Moldenke (loc. cit. 1956).  V. amaniensis and V. lokundjensis show some superficial similarities (both are tall rain forest trees with leaves drying dark, leaflets petiolulate (to 2.5 cm long) and flowers in lax few-flowered axillary compound dichasia).  However, this mainly Cameroon species differs in many characters from our plant: in the terete petiole, longer bracteoles, lanate calyx, glabrous and gland-dotted ovary and the eglandular hairs on filaments.  Our plant shows great uniformity and differs from many of the Tanzanian plants in the non-gland-dotted leaves, obsolete calyx teeth and median lower lobe of the corolla larger and yellow.  Some Tanzanian specimens (e.g. Ulugurue, Tanana, Morogoro, E.M. Bruce 774 (K)) approach our plants, being clearly intermediates of a range of variation.  Although there is a major disjunction between the Tanzanian and the Zimbabwean plants, both localities occur in similar montane habitats with a similar floristic composition.
Distribution
Zimbabwe District code: ZIM E, ZIM B, ZIM E, ZIM S, MOZ N, MOZ Z, MOZ T, MOZ MS, MOZ GI, MOZ M. Mozambique Also in Tanzania. Zambia
Morphology Reproductive morphology Inflorescences
Inflorescences axillary, 7–11 cm long, of ± slender, few-flowered compound dichasia, shortly crisped-pubescent; bracteoles 0.4–0.5 cm long, linear. Inflorescences axillary, 7–11 cm long, of ± slender, few-flowered compound dichasia, shortly crisped-pubescent; bracteoles 0.4–0.5 cm long, linear
Morphology Reproductive morphology Flowers Calyx
Calyx obconical, 5-toothed, not gland-dotted, densely pubescent, accrescent in fruit; tube c. 2 mm long; teeth c. 0.3 mm, erect. Calyx obconical, 5-toothed, not gland-dotted, densely pubescent, accrescent in fruit; tube c. 2 mm long; teeth c. 0.3 mm, erect
Morphology Reproductive morphology Flowers Corolla
Corolla pale yellow, 10–11 mm long, with scattered gland-dots, densely pubescent; tube c. 5 mm long, straight; lobes at obtuse angle to tube, middle lower lobe 5.5 × 4.5 cm. Corolla pale yellow, 10–11 mm long, with scattered gland-dots, densely pubescent; tube c. 5 mm long, straight; lobes at obtuse angle to tube, middle lower lobe 5.5 × 4.5 cm
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens scarcely exserted from tube; filaments pubescent at base, hairs eglandular. Stamens scarcely exserted from tube; filaments pubescent at base, hairs eglandular
Morphology Reproductive morphology Flowers Gynoecium Pistil
Ovary obovoid, sparsely gland-dotted at apex only, glabrous, truncate at apex; style 5 mm long.
Morphology Reproductive morphology Fruits
Drupe c. 1.4 × 1 cm, obovoid (?immature), rounded at apex, ± enclosed in the enlarged calyx; fruiting calyx cup-shaped, pubescent, not gland-dotted. Drupe c. 1.4 × 1 cm, obovoid (?immature), rounded at apex, ± enclosed in the enlarged calyx; fruiting calyx cup-shaped, pubescent, not gland-dotted.

[FTEA]

Verbenaceae, B. Verdcourt. Flora of Tropical East Africa. 1992

Morphology General Habit
Tree 12–20 m. tall, with smooth white bark or pale grey-brown ± corky bark on twigs; branchlets crisply pilose.
Morphology Leaves
Leaves drying dark, 5-foliolate; leaflets elliptic to obovate-elliptic, 6.5–18 cm. long, 3.2–9(–11) cm. wide, acuminate, cuneate at the base, glabrous above, pubescent beneath only on the nerves which are rather close and parallel, glandular-punctate, entire to remotely serrulate with upwardly directed teeth; petioles 6–12 cm. long; petiolules 0.5–2 cm. long.
Morphology Reproductive morphology Inflorescences
Inflorescences axillary, 8–25 cm. long, ± few-flowered dichasial cymes, the axes ferruginous tomentose or shortly crisped-pilose; bracteoles linear, 0.5–1 cm. long; peduncles 3–11.5(–13) cm. long; pedicels of central flowers up to 5 mm. long but others much shorter.
Morphology Reproductive morphology Flowers Calyx
Calyx campanulate, 3–4 mm. long, somewhat 2-lipped, densely grey crisped-hairy, irregularly toothed, the teeth ± 1 mm. long.
Morphology Reproductive morphology Flowers Corolla
Corolla lilac-white or pale blue with deeper blue lips and mauve lines; tube 3.5 mm. long, ferruginous tomentose outside; larger lip 3-fid, the main lobe 2.75 × 3 mm., the others much smaller; smaller lip with 3 ovate-triangular lobes 2.2 × 1–2 mm.
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary glandular at apex.
Morphology Reproductive morphology Fruits
Fruit black, ellipsoid-oblong, 0.9–1.1 cm. long, 6.5 mm. wide, glabrous; calycine cup 0.7–1 cm. wide, pale brown pubescent.
Habitat
Rain-forest, forest edges and relicts; 900–1400 m.
Distribution
not known elsewhere T3 T6 T7

Native to:

Tanzania, Zimbabwe

Vitex amaniensis W.Piep. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Braun [1054], Tanzania K000192778

First published in Beibl. Bot. Jahrb. Syst. 141: 60 (1928)

Accepted by

  • Fernandes, R. & Diniz, M.A. (2005). Avicenniaceae, Nesogenaceae, Verbenaceae and Lamiaceae (subfams, Viticoideae and Ajugoideae) Flora Zambesiaca 8(7): 1-161. Royal Botanic Gardens, Kew.
  • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

Literature

Flora Zambesiaca

  • Brenan, Check-list For. Trees Shrubs Tang. Terr: 643 (1949).
  • F.T.E.A., Verbenaceae: 58, 66 (1992).

Kew Backbone Distributions

  • Fernandes, R. & Diniz, M.A. (2005). Avicenniaceae, Nesogenaceae, Verbenaceae and Lamiaceae (subfams, Viticoideae and Ajugoideae) Flora Zambesiaca 8(7): 1-161. Royal Botanic Gardens, Kew.

Flora of Tropical East Africa

  • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 643 (1949).
  • Pieper in A. Engler, Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 62, Beibl. 141: 46, 60 (1928) (in key)
  • Pieper in F. Fedde, Repertorium Speciorum Novarum Regni Vegetabilis 26: 164 (1929).

Flora Zambesiaca
Flora Zambesiaca
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0