Isodon lophanthoides (Buch.-Ham. ex D.Don) H.Hara

This species is accepted, and its native range is Indian Subcontinent to China and Peninsula Malaysia.


Suddee, S., A. J. Paton, & Parnell, J. (2004). A Taxonomic Revision of Tribe Ocimeae Dumort. (Lamiaceae) in Continental South East Asia I. Introduction, Hyptidinae & Hanceolinae. Kew Bulletin, 59(3), 337-378. doi:10.2307/4110949

Morphology General Habit
Erect or ascending annual or short-lived perennial herb or undershrub up to 1.5 m tall
Morphology Leaves
Leaves well-spaced on the stem or basally clustered, membranous or chartaceous, broadly ovate, ovate, ovate-oblong or ovate-lanceolate, 15 - 150 x 8 - 80 mm, apex acute, obtuse or acuminate, base broadly cuneate, rounded, subtruncate or sometimes cordate, margin serrate or crenate, glabrescent, pubescent, scaberulous or scabrid above, glabrescent to pubescent and dotted with minute sessile glands beneath, greenish, purplish or yellowish-brown when dry, veins subprominent both sides; petioles up to 50 mm long in lower leaves, gradually reduced in length and becoming sessile on upper leaves, glabrescent to pubescent
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens long- exserted, villous at base; anterior pair inserted above the middle of corolla tube; posterior pair inserted near the base of corolla tube
Morphology Reproductive morphology Flowers Calyx
Calyx campanulate, 1 - 2 mm long at anthesis, glabrescent to densely pubescent with hairs denser at tube base, tubular-campanulate, 3 - 4 mm long in fruit, inconspicuously 2-lipped; teeth subequal, triangular- ovate, usually obtuse at apex; tube declinate, 10- nerved, twice as long as teeth, glabrescent to pubescent, with sessile glands outside, not gibbous at anterior base
Morphology Reproductive morphology Flowers Corolla
Corolla white, straight, 3 - 8 mm long; posterior lip subequally 4-lobed, lobes rounded or obtuse at apex, glabrous or pubescent, with sessile glands on back; anterior lip orbicular, 1 - 3 mm long, subequal to posterior, flattened or slightly concave, glabrous all over or slightly pubescent at base of lobe inside, glabrous or pubescent, with sessile glands outside; tube 2 - 5 mm long, subtubular, slightly dilated at throat, glabrous or slightly pubescent, with or without sessile glands outside
Morphology Reproductive morphology Flowers Disc
Disc annular with equal lobes or with anterior side well developed but not exceeding ovary
Morphology Reproductive morphology Flowers Gynoecium Style
Style long-exserted, longer than stamens
Morphology Reproductive morphology Fruits
Nutlets brown, ovoid or oblong, 0.8 - 1 mm long, smooth, producing mucilage when wet or not.
Morphology Reproductive morphology Inflorescences
Inflorescence terminal and axillary, composed of several lax narrow racemose branches, forming a large panicle, with axis glabrous to pubescent; cymes with peduncle up to 15 mm long, lax or dense with 3 - 18 flowers; bracts ovate, adaxially glabrous or glabrescent, abaxially glabrescent to pubescent, with sessile glands, persistent; bracteoles linear or linear-oblong, 0.5 - 2 mm long, pubescent, caducous or persistent; pedicels up to 8 mm long in flower and fruit, pubescent
Morphology Stem
Stems quadrangular, branched or not, glabrous to pubescent
So far, no specimen annotated by Hamilton as Hyssopus? lophanthoides from Narainhetty, Nepal has been seen. The specimens Bentham cited from Kumaon (Kamaon) (Wall. Cat. 2742B) are actually Isodon lophanthoides var. lophanthoides. All names based on P. gerardianus Benth. are also illegitimate. Wall. Cat. 2740, the type of Plectrantus striatus Benth. (Bentham 1830b) is also a syntype of P. graciliflorus Benth. (Bentham 1848). The name P. striatus was later applied by Bentham (1832, 1848), Hooker (1885) and several later authors to a form now recognised as I. lophanthoides var. lophanthoides. See under var. graciliflorus for more detail. Dunn (1913) recognised Bentham's Plectranthus gerardianus (nom. illeg.) and P. striatus in Wall. P1. Asiat. as the same species and mentioned that 'the types in Kew Herb. are connected by a continuous series of gradations with regard to their distinguishing characters'. In this study, with combination of other characters, the former is recognised under var. lophanthoides and the latter is recognised under var. graciliflorus. Isodon lophanthoides (Buch.-Ham. ex D. Don) H. Hara var. micranthus (C. Y. Wu) H. W. Li is the same as var. lophanthoides. Li (1988) mentioned that the corolla is 2 - 3 mm long and the leaves are purplish underneath when dry. The size of the corolla is not very different from var. lophanthoides and several specimens of var. lophanthoides from India, Thailand and Vietnam also have leaves which are purplish beneath when dry.
Nepal, Robert Blinkworth in Wall. Cat. 2742B (K-W! neotype chosen here; isoneotypes BM!, K!).

Native to:

Assam, Bangladesh, China North-Central, China South-Central, China Southeast, East Himalaya, India, Laos, Malaya, Myanmar, Nepal, Thailand, Tibet, Vietnam, West Himalaya

Isodon lophanthoides (Buch.-Ham. ex D.Don) H.Hara appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Jan 5, 2017 Nguyen, V.D. [HNK-2133(RK 1244)], Vietnam K000249938
Jan 1, 2014 Staples [1462], Thailand K000938613
Jun 1, 2013 Ducloux [7331], China K000938063
Jun 1, 2013 Delavay [s.n.], China K000938064
Jun 1, 2013 Ducloux [5785], China K000938065
Jun 1, 2013 Bodinier [2480], China K000938066
Jun 1, 2013 Bodinier [2852], China K000938067
Jun 1, 2013 Poilane [28378], Vietnam K000938078
Jan 1, 2010 Du, al. [HNK 2182], Vietnam K000249669
Jan 1, 2009 Santisuk [1523], Thailand K000248477
Jan 1, 2009 Bai-Zhong [4825], China K000248764
Jan 1, 2001 Benson, R. [1166], Myanmar K001067871 syntype
Jan 1, 2001 Griffith, W. [214], India K001067857
Jan 1, 2001 Wallich, N. [2742], India K001067858 neotype
Nguyen, V.D. [HNK-2138], Vietnam K000249937
Clark, C.B. [12678], India K001067864
Heyne, B. [s.n.], India K001067870 syntype
Stocks, J.E. [s.n.], India K001067866 syntype
Clark, C.B. [25271], India K001067862
Hooker, J.D. [s.n.], India K001067860
Stocks, J.E. [s.n.], India K001067868 syntype
Clark, C.B. [25257], India K001067861
Stocks, J.E. [s.n.], India K001067865 syntype
s.coll. [2740], Bangladesh K001067856
Stocks, J.E. [s.n.], India K001067869 syntype
Stocks, J.E. [s.n.], India K001067867 syntype
Hooker, J.D. [s.n.], India K001067863

First published in J. Jap. Bot. 60: 235 (1985)

Accepted by

  • Bramley, G.L.C. (2019). Flora Malesiana 23: 1-444. Noordhoff-Kolff N.V., Djakarta.
  • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
  • Khanam, M. & Hassan, M.A. (2008). Lamiaceae Flora of Bangladesh 58: 1-161. Bangladesh National Herbarium, Dhaka.


Kew Backbone Distributions

  • Bramley, G.L.C. (2019). Flora Malesiana 23: 1-444. Noordhoff-Kolff N.V., Djakarta.

  • Art and Illustrations in Digifolia

    Digital Image © Board of Trustees, RBG Kew

  • Herbarium Catalogue Specimens

    Digital Image © Board of Trustees, RBG Kew

  • Kew Backbone Distributions

    The International Plant Names Index and World Checklist of Selected Plant Families 2022. Published on the Internet at and
    © Copyright 2017 World Checklist of Selected Plant Families.

  • Kew Bulletin

    Kew Bulletin

  • Kew Names and Taxonomic Backbone

    The International Plant Names Index and World Checklist of Selected Plant Families 2022. Published on the Internet at and
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.