Phyllanthus tenellus Roxb.

Native to:

Comoros, Madagascar, Mauritius, Mozambique, Réunion, Saudi Arabia, Tanzania, Yemen

Introduced into:

Alabama, Angola, Argentina Northeast, Azores, Bahamas, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Canary Is., Cape Verde, Caroline Is., Cayman Is., Cook Is., Costa Rica, Florida, Georgia, Hawaii, Illinois, India, Iran, Jamaica, Japan, Leeward Is., Louisiana, Madeira, Marquesas, Mexico Central, Mexico Southwest, Mississippi, Nansei-shoto, New Caledonia, New Guinea, New South Wales, Norfolk Is., North Carolina, Northern Territory, Puerto Rico, Queensland, Seychelles, Sicilia, Society Is., South Carolina, Sri Lanka, Taiwan, Tennessee, Texas, Tuamotu, Tubuai Is., Vanuatu, Venezuela, Virginia, Western Australia, Windward Is.

POWO follows these authorities in accepting this name:

• Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies. Smithsonian Contributions to Botany 98: 1-1192.
• Bailey, C. & al. (2015). Guide to the Vascular Plants of Tennessee: 1-813. University of Tennessee press.
• Balakrishnan, N.P. & Chakrabarty, T. (2007). The family Euphorbiaceae in India. A synopsis of its profile, taxonomy and bibliography: 1-500. Bishen Singh Mahendra Pal Singh, Dehra Dun.
• Chakrabarty, T. & Balakrishnan, N.P. (2018). Indo-Burmese Phyllanthaceae: A Taxonomic Revision: 1-437. Bishen Singh Mahendra Pal Singh, Dehra Dun.
• Crisafulli, A., Picone, R.M. & Zaccone, S. (2011). Phyllanthus tenellus (Phyllanthaceae) a new alien species naturalised to Sicily, first record for Italy. Flora Mediterranea 21: 293-297.
• Dobignard, A. & Chatelain, C. (2013). Index synonymique de la flore d'Afrique du nord 5: 1-451. Éditions des conservatoire et jardin botaniques, Genève.
• Figueiredo, E. & Smith, G.F. (2008). Plants of Angola. Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
• Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1-1622. The Board of Trustees of the Royal Botanic Gardens, Kew.
• Hammel, B.E., Grayum, M.H., Herrera & C. & Zamora, N. (eds.) (2010). Manual de plantas de Costa Rica volumen V. Dicotiledóneas (Clusiaceae-Gunneraceae). Monographs in Systematic Botany from the Missouri Botanical Garden 119: 1-970. Missouri Botanical Garden.
• Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
• Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
• Kuo, M.L. (ed.) (2012). Flora of Taiwan, ed. 2, Suppl.: 1-414. Editorial Committee of the Flora of Taiwan, Second Edition, National Taiwan University.
• MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
• Martins, E.R. & de Lima, L.R. (2011). Sinopse do gênero Phyllanthus L. (Phyllanthaceae) do estado de São Paulo. Hoehnea 38: 123-133.
• Mohlenbrock, R.H. (2014). Vascular Flora of Illinois. A Field Guide, ed. 4: 1-536. Southern Illinois University Press, Carbondale.
• Muer, T., Sauerbier, H. & Cabrara Calixto, F. (2020). Die Farn- und Blütenpflanzen Madeiras: 1-792. Verlag und Versandbuchhandlung Andreas Kleinsteuber.
• Orchard, A.E. (ed.) (1994). Oceanic Islands 1. Flora of Australia 49: 1-681. Australian Government Publishing Service, Canberra.
• Plunkett, G.M., Ranker, T.A., Sam, C. & Balick, M.J. (2022). Towards a checklist of the vascular flora of Vanuatu. Candollea 77: 105-118.
• Ralimanana, H. & Hoffmann, P. (2011 publ. 2012). Taxonomic revision of Phyllanthus (Phyllanthaceae) in Madagascar and the Comoro Islands: synopsis and subgenera Isocladus, Betsileani, Kirganelia and Tenellanthus. Kew Bulletin 66: 331-365.
• Schäfer, H. (2021). Flora of the Azores a field guide: 1-445. Margraf Publishers GmbH.
• Sykes, W.R. (2016). Flora of the Cook Islands: 1-973. National Tropical Botanical Garden, Hawaii.
• Zuloaga, F.O. & Belgrano, M.J. (eds.) (2017). Flora Argentina. Flora vascular de la República Argentina 17: 1-434. INTA, IMBIV & IBODA.
• da Silva, M.J. & de Sales, M.F. (2008). Sinopse do Gênero Phyllanthus (Phyllanthaceae) no Nordeste do Brasil. Rodriguésia; Revista do Instituto de Biologia Vegetal, Jardim Botânico e Estaçao Biologica do Itatiaya 59: 401-406.

Flora of the Cayman Islands

• Roxb., Fl. Ind. 3: 668 (1832).

Kew Bulletin

• (Roxburgh 1832: 668).
• Baillon, H. (1858). Étude générale du groupe des Euphorbiacées. Victor Masson, Paris.
• Baillon, H. (1861). Recueil périodique d’observations botaniques. Adansonia 2: 47 – 51.
• Brunel, J. F. & Roux, J. (1981). Phyllantheae de Madagascar II: à propos du complexe Phyllanthus nummulariaefolius [sic] Poir. — Phyllanthus tenellus Roxb. Bull. Mus. Hist. Nat., Paris, série 4, 3, section B, Adansonia 2: 185 – 199.
• Brunel, J. F. (1987). Sur le genre Phyllanthus L. et quelques genres voisins de la Tribu des Phyllantheae Dumort. (Euphorbiaceae, Phyllantheae) en Afrique intertropicale et à Madagascar. Thèse doctorale, Université Louis Pasteur, Strasbourg.
• De Candolle, A. (1901). Plantae Madagascarienses ab Alberto Mocquerysio lectae. Bull. Herb. Boissier II, 1: 549 – 587.
• Dorr, L. J. (1997). Plant Collectors in Madagascar and the Comoro Islands. Royal Botanic Gardens, Kew.
• Friedmann, F. (1994). Flore des Seychelles. Editions de l’ORSTOM, Paris.
• Gmelin, J. F. (1792). Systema Naturae, ed. 13, vol. 2 (2). G. E. Beer, Leipzig.
• Govaerts, R., Frodin, D. G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (with Pandaceae), vols 1 – 4. Royal Botanic Gardens, Kew.
• Hoffmann, P. & McPherson, G. (2003). Transfer of Madagascan Glochidion to Phyllanthus. Novon 13: 307 – 310.CrossRefGoogle Scholar
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• Humboldt, F. W. H. von, Bonpland, A. J. A. & Kunth, C. S. (1817). Nova genera et species plantarum 2. Sumptibus Librairie Graeco-Latini-Germanicae, Paris.
• IUCN (2001). IUCN Red List Categories and Criteria: Version 3.1. IUCN Species Survival Commission, IUCN, Gland and Cambridge.
• Jussieu, A. L. de (1789). Genera Plantarum. Herissant & Barrois, Paris.
• Jussieu, A. L. de (1824). De Euphorbiacearum Generibus Medicisque Earumdem Viribus Tentamen. Didot, Paris.
• Kathriarachchi, H., Hoffmann, P., Samuel, R., Wurdack, K. J. & Chase, M. W. (2005). Molecular phylogenetics of Phyllanthaceae inferred from 5 genes (plastid atpB, matK, 3’ ndhF, rbcL) and nuclear PHYC. Molec. Phylogen. Evol. 36: 112 – 134.CrossRefGoogle Scholar
• Kathriarachchi, H., Samuel, R., Hoffmann, P., Mlinarec, J., Wurdack, K. J., Ralimanana, H., Stuessy, T. F. & Chase, M. W. (2006). Phylogenetics of tribe Phyllantheae (Phyllanthaceae; Euphorbiaceae sensu lato) based on nrITS and plastid matK DNA sequence data. Amer. J. Bot. 93: 637 – 655.CrossRefGoogle Scholar
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• Leandri, J. (1934). Espèces et localités nouvelles d’Euphorbiacées d’Afrique et Madagascar. Bull. Soc. Bot. France 81: 449 – 454.
• Leandri, J. (1938). Contribution à l’étude des Euphorbiacées de Madagascar (suite). Notul. Syst. (Paris) 6: 185 – 199.
• Leandri, J. (1939). Contribution à l’étude des Euphorbiacées de Madagascar. Notul. Syst. (Paris) 7: 168 – 187.
• Leandri, J. (1957). Notes systématiques sur les Euphorbiacées, Phyllanthus de Madagascar. Mém. Inst. Sci. Madagascar 8: 205 – 234.
• Leandri, J. (1958). Euphorbiaceae I (Phyllanthoideae). In: H. Humbert (ed.), Flore de Madagascar et des Comores, 111ème famille 1: 1 – 199. Firmin-Didot, Paris.
• Linnaeus, C. (1753). Species Plantarum, ed. 1, 2. Impensis Laurentii Salvii, Stockholm.
• McNeill, J., Barrie, F. R., Burdet, H. M., Demoulin, V., Hawksworth, D. L., Marhold, K., Nicolson, D. H., Prado, J., Silva, P. C., Skog, J. E., Wiersema, J. H. & Turland, N. J. (2006). International Code of Botanical Nomenclature. Adopted by the Seventeenth International Botanical Congress Vienna, Austria, July 2005. Regnum Veg. 146: 1 – 568.
• Müller, J. (1863). Euphorbiaceae. Vorläufige Mittheilungen aus dem für De Candolle’s Prodromus bestimmten Manuscript über diese Familie. Linnaea 32: 1 – 126.
• Müller, J. (1866). Euphorbiaceae. In: A. De Candolle (ed.), Prodromus Systematis Naturalis Regni Vegetabilis 15(2), pp. 189 – 1286. Victor Masson, Paris.
• Persoon, C. H. (1807). Synopsis Plantarum 2. C. F. Cramer, Paris & J. G. Cotta, Tübingen.
• Radcliffe-Smith, A. (1985). Notes on African Euphorbiaceae: XVI. Kew Bull. 40: 657 – 658.CrossRefGoogle Scholar
• Roxburgh, W. (1832). Flora Indica, ed. 2, vol. 3. W. Thacker & Co., Calcutta & Parbury, Allan & Co., London.
• Samuel, R., Kathriarachchi, H., Hoffmann, P., Barfuss, M., Wurdack, K. J. & Chase, M. W. (2005). Molecular phylogeny of Phyllanthaceae: Evidence from plastid matK and nuclear PHYC sequences. Amer. J. Bot. 92: 132 – 141.CrossRefGoogle Scholar
• Stafleu, F. A. & Cowan, R. S. (1976). Taxonomic Literature I: A – G. Regnum Veg. 94: 1 – 1136. Bohn, Scheltema & Holkema, Utrecht.
• Webster, G. L. & Miller K. L. (1963). The genus Reverchonia (Euphorbiaceae). Rhodora 65: 193 – 207.
• Webster, G. L. (1955). Studies of the Euphorbiaceae, Phyllanthoideae I. Taxonomic notes on the West Indian species of Phyllanthus. Contr. Gray Herb. Harv. Univ. 176: 45 – 63.
• Webster, G. L. (1956 – 1958). A monographic study of the West Indian species of Phyllanthus. J. Arnold Arbor. 37: 91 – 122, 217 – 268, 340 – 359; 38: 51 – 80, 170 – 198, 295 – 373; 39: 49 – 100, 111 – 212.
• Webster, G. L. (1967). The genera of the Euphorbiaceae in the Southeastern United States. J. Arnold Arbor. 48: 303 – 431.
• Willdenow, C. L. (1805). Species plantarum, ed. 4, vol. 4.1. G. C. Nauk, Berlin.
• Willemet, P. (1796). Herbarium Mauritianum. Ann. Bot. (Usteri) 18: 1 – 66.
• Wurdack, K. J., Hoffmann, P., Samuel, R., de Bruijn, A., van der Bank, M. & Chase, M. W. (2004). Molecular phylogenetic analysis of Phyllanthaceae (Phyllanthoideae pro parte, Euphorbiaceae sensu lato) using plastid rbcL DNA sequences. Amer. J. Bot. 91: 1882 – 1990.CrossRefGoogle Scholar

Kew Backbone Distributions

• (2022). https://doi.org/10.11646/phytotaxa.539.2.3. epublication.
• Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies. Smithsonian Contributions to Botany 98: 1-1192.
• Bailey, C. & al. (2015). Guide to the Vascular Plants of Tennessee: 1-813. University of Tennessee press.
• Balakrishnan, N.P. & Chakrabarty, T. (2007). The family Euphorbiaceae in India. A synopsis of its profile, taxonomy and bibliography: 1-500. Bishen Singh Mahendra Pal Singh, Dehra Dun.
• Crisafulli, A., Picone, R.M. & Zaccone, S. (2011). Phyllanthus tenellus (Phyllanthaceae) a new alien species naturalised to Sicily, first record for Italy. Flora Mediterranea 21: 293-297.
• Flora of North America Editorial Committee (2016). Flora of North America North of Mexico 12: 1-603. Oxford University Press, New York, Oxford.
• Florence, J. (1997). Flore de la Polynésie Française 1: 1-393. ORSTOM éditions, Paris.
• Fosberg, F.R., Sachet, M.-H., Oliver, R. (1979). A geographical checklist of the Micronesian Dicotyledonae. Micronesica; Journal of the College of Guam 15: 41-295.
• Hammel, B.E., Grayum, M.H., Herrera & C. & Zamora, N. (eds.) (2010). Manual de plantas de Costa Rica volumen V. Dicotiledóneas (Clusiaceae-Gunneraceae). Monographs in Systematic Botany from the Missouri Botanical Garden 119: 1-970. Missouri Botanical Garden.
• Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
• MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
• Martins, E.R. & de Lima, L.R. (2011). Sinopse do gênero Phyllanthus L. (Phyllanthaceae) do estado de São Paulo. Hoehnea 38: 123-133.
• Mohlenbrock, R.H. (2014). Vascular Flora of Illinois. A Field Guide, ed. 4: 1-536. Southern Illinois University Press, Carbondale.
• Muer, T., Sauerbier, H. & Cabrara Calixto, F. (2020). Die Farn- und Blütenpflanzen Madeiras: 1-792. Verlag und Versandbuchhandlung Andreas Kleinsteuber.
• Orchard, A.E. (ed.) (1994). Oceanic Islands 1. Flora of Australia 49: 1-681. Australian Government Publishing Service, Canberra.
• Ralimanana, H. & Hoffmann, P. (2011 publ. 2012). Taxonomic revision of Phyllanthus (Phyllanthaceae) in Madagascar and the Comoro Islands: synopsis and subgenera Isocladus, Betsileani, Kirganelia and Tenellanthus. Kew Bulletin 66: 331-365.
• Sykes, W.R. (2016). Flora of the Cook Islands: 1-973. National Tropical Botanical Garden, Hawaii.
• da Silva, M.J. & de Sales, M.F. (2008). Sinopse do Gênero Phyllanthus (Phyllanthaceae) no Nordeste do Brasil. Rodriguésia; Revista do Instituto de Biologia Vegetal, Jardim Botânico e Estaçao Biologica do Itatiaya 59: 401-406.